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| Lichtig, Asher J., Lucas, Spencer G. and Alvarado, Guillermo E. Reappraisal of the holotype of “testudo” costarricensis, a tortoise supposedly from the oligocene or lower miocene of Costa Rica. Rev. Geol. Amér. Central, Dec 2018, no.59, p.39-47. ISSN 0256-7024
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