<?xml version="1.0" encoding="ISO-8859-1"?><article xmlns:mml="http://www.w3.org/1998/Math/MathML" xmlns:xlink="http://www.w3.org/1999/xlink" xmlns:xsi="http://www.w3.org/2001/XMLSchema-instance">
<front>
<journal-meta>
<journal-id>0034-7744</journal-id>
<journal-title><![CDATA[Revista de Biología Tropical]]></journal-title>
<abbrev-journal-title><![CDATA[Rev. biol. trop]]></abbrev-journal-title>
<issn>0034-7744</issn>
<publisher>
<publisher-name><![CDATA[Universidad de Costa Rica]]></publisher-name>
</publisher>
</journal-meta>
<article-meta>
<article-id>S0034-77442021000600304</article-id>
<article-id pub-id-type="doi">10.15517/rbt.v69is2.48745</article-id>
<title-group>
<article-title xml:lang="es"><![CDATA[Comportamiento diurno de cetáceos en el Área de Conservación Guanacaste, Costa Rica]]></article-title>
<article-title xml:lang="en"><![CDATA[Cetacean diurnal behavior in the Guanacaste Conservation Area, Costa Rica]]></article-title>
</title-group>
<contrib-group>
<contrib contrib-type="author">
<name>
<surname><![CDATA[Martínez-Fernández]]></surname>
<given-names><![CDATA[Damián]]></given-names>
</name>
<xref ref-type="aff" rid="Aff"/>
</contrib>
</contrib-group>
<aff id="Af1">
<institution><![CDATA[,Federación Costarricense de Pesca Director del Departamento de Política y Conservación ]]></institution>
<addr-line><![CDATA[ San José]]></addr-line>
<country>Costa Rica</country>
</aff>
<pub-date pub-type="pub">
<day>00</day>
<month>10</month>
<year>2021</year>
</pub-date>
<pub-date pub-type="epub">
<day>00</day>
<month>10</month>
<year>2021</year>
</pub-date>
<volume>69</volume>
<fpage>304</fpage>
<lpage>312</lpage>
<copyright-statement/>
<copyright-year/>
<self-uri xlink:href="http://www.scielo.sa.cr/scielo.php?script=sci_arttext&amp;pid=S0034-77442021000600304&amp;lng=en&amp;nrm=iso"></self-uri><self-uri xlink:href="http://www.scielo.sa.cr/scielo.php?script=sci_abstract&amp;pid=S0034-77442021000600304&amp;lng=en&amp;nrm=iso"></self-uri><self-uri xlink:href="http://www.scielo.sa.cr/scielo.php?script=sci_pdf&amp;pid=S0034-77442021000600304&amp;lng=en&amp;nrm=iso"></self-uri><abstract abstract-type="short" xml:lang="es"><p><![CDATA[Resumen  Introducción:  Tres especies de cetáceos, el delfín manchado pantropical (Stenella attenuata attenuata), el delfín nariz de botella (Tursiops truncatus) y la ballena jorobada (Megaptera novaeangliae) son un componente importante de la fauna marina del Área de Conservación Guanacaste. Sin embargo, la información biológica básica sobre el comportamiento y la presencia de estas especies sigue siendo escasa.  Objetivo:  El objetivo de este estudio es determinar el comportamiento diurno de los cetáceos en el sector marino de la ACG y en sus áreas adyacentes.  Métodos:  Se muestreó el Pacífico norte de Costa Rica desde Bahía Salinas hasta las Islas Murciélago, incluyendo el Golfo de Santa Elena, entre mayo de 2005 y mayo de 2006. Se utilizó el método de escaneo de comportamiento para identificar el comportamiento general de los grupos.  Resultados:  Se registraron encuentros para tres especies en un total de 36 avistamientos, pero solo con un tiempo considerable para dos especies, S. attenuata y M. novaeangliae. Los delfines manchados exhibieron una amplia variedad de comportamientos en la temporada de lluvias. La búsqueda de alimento (X2 = 29.42; g.l. = 1; P &lt; 0.001) y los viajes (X2 = 38.33; g.l. = 1; P &lt; 0.001) aumentaron significativamente en la época de lluvias. Los comportamientos predominantes para la ballena jorobada fueron la asociación madre-cría, social y de viaje; este último fue mayor en la época de lluvias (X2 = 78.16; g.l. = 1; P &lt; 0.001).  Conclusiones:  En la zona, los cetáceos presentaron comportamientos como forrajeo y madre-cría, lo que significa que el área puede clasificarse como un hábitat crítico para estas especies de cetáceos, de mayor importancia para especies altamente migratorias como las ballenas jorobadas.]]></p></abstract>
<abstract abstract-type="short" xml:lang="en"><p><![CDATA[Abstract  Introduction:  Three species of cetaceans, the pantropical spotted dolphin (Stenella attenuata attenuata), the bottlenose dolphin (Tursiops truncatus) and the humpback whale (Megaptera novaeangliae), are important components of the marine fauna of the Guanacaste Conservation Area (ACG). However, basic biological information on the behavior and occurrence of these species remains scarce.  Objective:  The objective of this study is to determine the diurnal behavior of the three-mentioned species in the ACG.  Methods:  The north Pacific of Costa Rica, from Bahía Salinas to the Marine Sector of ACG was sampled between May 2005 and May 2006. The behavior scanning method was used to identify the general behavior of the cetacean groups.  Results:  Encounters were recorded for the three species, in a total of 36 sightings, but only for S. attenuata and M. novaeangliae for considerable periods of time. Stenella attenuata exhibited a wide variety of behaviors, being foraging (X2 = 29.42; d.f. = 1; P &lt; 0.001) and travelling (X2 = 38.33; d.f. = 1; P &lt; 0.001) significantly higher during the rainy season. Predominant behaviors for the humpback whale were mother-calf association, social and travelling; being the latter higher during the rainy season (X2 = 78.16; d.f. = 1; P &lt; 0.001).  Conclusions:  Behaviors such as foraging and mother-calf were the most representative among species and the most frequent. This suggests that the area can be considered as a critical habitat for these species of cetaceans, with particular importance for highly migratory species such as humpback whales.]]></p></abstract>
<kwd-group>
<kwd lng="en"><![CDATA[dolphins]]></kwd>
<kwd lng="en"><![CDATA[humpback whales]]></kwd>
<kwd lng="en"><![CDATA[Stenella attenuata]]></kwd>
<kwd lng="en"><![CDATA[Megaptera novaeangliae]]></kwd>
<kwd lng="en"><![CDATA[cetacean behavior]]></kwd>
<kwd lng="en"><![CDATA[mother-calf association]]></kwd>
<kwd lng="en"><![CDATA[marine conservation]]></kwd>
<kwd lng="es"><![CDATA[delfines]]></kwd>
<kwd lng="es"><![CDATA[ballenas jorobadas]]></kwd>
<kwd lng="es"><![CDATA[Stenella attenuata]]></kwd>
<kwd lng="es"><![CDATA[Megaptera novaeangliae]]></kwd>
<kwd lng="es"><![CDATA[comportamiento de cetáceos]]></kwd>
<kwd lng="es"><![CDATA[conservación marina]]></kwd>
</kwd-group>
</article-meta>
</front><back>
<ref-list>
<ref id="B1">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Acevedo]]></surname>
<given-names><![CDATA[A.]]></given-names>
</name>
<name>
<surname><![CDATA[Smultea]]></surname>
<given-names><![CDATA[M. A.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[First records of humpback whales including calves at Golfo Dulce and Isla del Coco, Costa Rica, suggesting geographical overlap of Northern and Southern Hemisphere populations]]></article-title>
<source><![CDATA[Marine Mammal Science]]></source>
<year>1995</year>
<volume>11</volume>
<numero>4</numero>
<issue>4</issue>
<page-range>554-60</page-range></nlm-citation>
</ref>
<ref id="B2">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Acevedo]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
<name>
<surname><![CDATA[Aguayo-Lobo]]></surname>
<given-names><![CDATA[A.]]></given-names>
</name>
<name>
<surname><![CDATA[Allen]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
<name>
<surname><![CDATA[Botero-Acosta]]></surname>
<given-names><![CDATA[N.]]></given-names>
</name>
<name>
<surname><![CDATA[Capella]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
<name>
<surname><![CDATA[Castro]]></surname>
<given-names><![CDATA[C.]]></given-names>
</name>
<name>
<surname><![CDATA[Stevick]]></surname>
<given-names><![CDATA[P. T.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Migratory preferences of Humpback Whales between feeding and breeding grounds in the Eastern South Pacific]]></article-title>
<source><![CDATA[Marine Mammal Science]]></source>
<year>2017</year>
<volume>33</volume>
<numero>4</numero>
<issue>4</issue>
<page-range>1035-52</page-range></nlm-citation>
</ref>
<ref id="B3">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Alfaro]]></surname>
<given-names><![CDATA[E. J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Caracterización del &#8216;Veranillo&#8217; en dos cuencas de la Vertiente del Pacífico de Costa Rica, América Central]]></article-title>
<source><![CDATA[Revista de Biología Tropical]]></source>
<year>2014</year>
<volume>62</volume>
<numero>^s4</numero>
<issue>^s4</issue>
<supplement>4</supplement>
<page-range>S1-S15</page-range></nlm-citation>
</ref>
<ref id="B4">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Alfaro]]></surname>
<given-names><![CDATA[E. J.]]></given-names>
</name>
<name>
<surname><![CDATA[Cortés]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Atmospheric forcing of cool and subsurface water events in Bahía Culebra, Gulf of Papagayo, Costa Rica]]></article-title>
<source><![CDATA[Revista de Biología Tropcical]]></source>
<year>2012</year>
<volume>60</volume>
<numero>^s2</numero>
<issue>^s2</issue>
<supplement>2</supplement>
<page-range>S173-86</page-range></nlm-citation>
</ref>
<ref id="B5">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Alfaro]]></surname>
<given-names><![CDATA[E. J.]]></given-names>
</name>
<name>
<surname><![CDATA[Cortés]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Forcing of cool and warm subsurface water events in Bahía Salinas, Costa Rica]]></article-title>
<source><![CDATA[Revista de Biología Tropical]]></source>
<year>2021</year>
<volume>69</volume>
<numero>^s2</numero>
<issue>^s2</issue>
<supplement>2</supplement>
<page-range>S127-41</page-range></nlm-citation>
</ref>
<ref id="B6">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Alfaro]]></surname>
<given-names><![CDATA[E. J.]]></given-names>
</name>
<name>
<surname><![CDATA[Cortés]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
<name>
<surname><![CDATA[Alvarado]]></surname>
<given-names><![CDATA[J. J.]]></given-names>
</name>
<name>
<surname><![CDATA[Jiménez]]></surname>
<given-names><![CDATA[C.]]></given-names>
</name>
<name>
<surname><![CDATA[León]]></surname>
<given-names><![CDATA[A.]]></given-names>
</name>
<name>
<surname><![CDATA[Sánchez-Noguera]]></surname>
<given-names><![CDATA[C.]]></given-names>
</name>
<name>
<surname><![CDATA[Nivia-Ruiz]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
<name>
<surname><![CDATA[Ruiz]]></surname>
<given-names><![CDATA[E.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Clima y temperatura sub-superficial del mar en Bahía Culebra, Golfo de Papagayo, Costa Rica]]></article-title>
<source><![CDATA[Revista de Biología Tropical]]></source>
<year>2012</year>
<volume>60</volume>
<numero>^s2</numero>
<issue>^s2</issue>
<supplement>2</supplement>
<page-range>S159-71</page-range></nlm-citation>
</ref>
<ref id="B7">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Altmann]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Observational study of behavior: bampling methods]]></article-title>
<source><![CDATA[Behaviour]]></source>
<year>1974</year>
<volume>49</volume>
<numero>3-4</numero>
<issue>3-4</issue>
<page-range>227-66</page-range></nlm-citation>
</ref>
<ref id="B8">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Baird]]></surname>
<given-names><![CDATA[R. W.]]></given-names>
</name>
<name>
<surname><![CDATA[Dill]]></surname>
<given-names><![CDATA[L. M.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Occurrence and behaviour of transient Killer Whales: seasonal and pod-specific variability, foraging behaviour, and prey handling]]></article-title>
<source><![CDATA[Canadian Journal of Zoology]]></source>
<year>1995</year>
<volume>73</volume>
<numero>7</numero>
<issue>7</issue>
<page-range>1300-11</page-range></nlm-citation>
</ref>
<ref id="B9">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Baird]]></surname>
<given-names><![CDATA[R. W.]]></given-names>
</name>
<name>
<surname><![CDATA[Ligon]]></surname>
<given-names><![CDATA[A. D.]]></given-names>
</name>
<name>
<surname><![CDATA[Hooker]]></surname>
<given-names><![CDATA[S. K.]]></given-names>
</name>
<name>
<surname><![CDATA[Gorgone]]></surname>
<given-names><![CDATA[A. M.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Subsurface and nighttime behavior of Pantropical Spotted Dolphins in Hawaii]]></article-title>
<source><![CDATA[Canadian Journal of Zoology]]></source>
<year>2001</year>
<volume>79</volume>
<page-range>988-96</page-range></nlm-citation>
</ref>
<ref id="B10">
<nlm-citation citation-type="book">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Bassey]]></surname>
<given-names><![CDATA[F. G.]]></given-names>
</name>
</person-group>
<source><![CDATA[Evaluación ecológica de los arrecifes y comunidades coralinas de las Islas Murciélago y sección norte de la Península de Santa Elena, Pacífico de Costa Rica]]></source>
<year>2010</year>
<publisher-loc><![CDATA[Puntarenas, Costa Rica ]]></publisher-loc>
<publisher-name><![CDATA[Universidad Nacional de Costa Rica]]></publisher-name>
</nlm-citation>
</ref>
<ref id="B11">
<nlm-citation citation-type="book">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Bettridge]]></surname>
<given-names><![CDATA[S. C.]]></given-names>
</name>
<name>
<surname><![CDATA[Baker]]></surname>
<given-names><![CDATA[S.]]></given-names>
</name>
<name>
<surname><![CDATA[Barlow]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
<name>
<surname><![CDATA[Clapham]]></surname>
<given-names><![CDATA[P. J.]]></given-names>
</name>
<name>
<surname><![CDATA[Ford]]></surname>
<given-names><![CDATA[M.]]></given-names>
</name>
<name>
<surname><![CDATA[Gouveia]]></surname>
<given-names><![CDATA[D.]]></given-names>
</name>
<name>
<surname><![CDATA[Wade]]></surname>
<given-names><![CDATA[P. R.]]></given-names>
</name>
</person-group>
<source><![CDATA[Status review of the Humpback Whale (Megaptera novaeangliae) under the Endangered Species Act]]></source>
<year>2015</year>
<publisher-name><![CDATA[Department of Commerce, NOAA Technical Memorandum NMFS NOAA-TMNMFS-SWFSC-540. National Oceanic and Atmospheric Administration, National Marine Fisheries Service, Southwest Fisheries Science Center, La Jolla]]></publisher-name>
</nlm-citation>
</ref>
<ref id="B12">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Bianchi]]></surname>
<given-names><![CDATA[G.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Demersal assemblages of the continental shelf and slope edge between the Gulf of Tehuantepec (Mexico) and the Gulf of Papagayo (Costa Rica)]]></article-title>
<source><![CDATA[Marine Ecology Progress Series]]></source>
<year>1991</year>
<volume>73</volume>
<numero>2-3</numero>
<issue>2-3</issue>
<page-range>121-40</page-range></nlm-citation>
</ref>
<ref id="B13">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Burt]]></surname>
<given-names><![CDATA[W. H.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Territoriality and home range concepts as applied to mammals]]></article-title>
<source><![CDATA[Journal of Mammalogy]]></source>
<year>1943</year>
<volume>24</volume>
<numero>3</numero>
<issue>3</issue>
<page-range>346-52</page-range></nlm-citation>
</ref>
<ref id="B14">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Calambokidis]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
<name>
<surname><![CDATA[Steiger]]></surname>
<given-names><![CDATA[G. H.]]></given-names>
</name>
<name>
<surname><![CDATA[Rasmussen]]></surname>
<given-names><![CDATA[K.]]></given-names>
</name>
<name>
<surname><![CDATA[Urbán]]></surname>
<given-names><![CDATA[J. R.]]></given-names>
</name>
<name>
<surname><![CDATA[Balcomb]]></surname>
<given-names><![CDATA[K. C.]]></given-names>
</name>
<name>
<surname><![CDATA[Ladrón De Guevara]]></surname>
<given-names><![CDATA[P. P.]]></given-names>
</name>
<name>
<surname><![CDATA[Darling]]></surname>
<given-names><![CDATA[J. D.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Migratory destinations of Humpback Whales that feed off California, Oregon and Washington]]></article-title>
<source><![CDATA[Marine Ecology Progress Series]]></source>
<year>2000</year>
<volume>192</volume>
<page-range>295-304</page-range></nlm-citation>
</ref>
<ref id="B15">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Calambokidis]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
<name>
<surname><![CDATA[Steiger]]></surname>
<given-names><![CDATA[G. H.]]></given-names>
</name>
<name>
<surname><![CDATA[Straley]]></surname>
<given-names><![CDATA[J. M.]]></given-names>
</name>
<name>
<surname><![CDATA[Herman]]></surname>
<given-names><![CDATA[L. M.]]></given-names>
</name>
<name>
<surname><![CDATA[Cerchio]]></surname>
<given-names><![CDATA[S.]]></given-names>
</name>
<name>
<surname><![CDATA[Salden]]></surname>
<given-names><![CDATA[D. R.]]></given-names>
</name>
<name>
<surname><![CDATA[Quinn]]></surname>
<given-names><![CDATA[T. J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Movements and population structure of Humpback Whales in the North Pacific]]></article-title>
<source><![CDATA[Marine Mammal Science]]></source>
<year>2001</year>
<volume>17</volume>
<numero>4</numero>
<issue>4</issue>
<page-range>769-94</page-range></nlm-citation>
</ref>
<ref id="B16">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Cortés]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Comunidades coralinas y arrecifes del Área de Conservación Guanacaste, Costa Rica]]></article-title>
<source><![CDATA[Revista de Biología Tropical]]></source>
<year>1996</year>
<volume>44</volume>
<numero>3</numero>
<issue>3</issue>
<page-range>623-5</page-range></nlm-citation>
</ref>
<ref id="B17">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Cortés]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Marine biodiversity baseline for Área de Conservación Guanacaste, Costa Rica: published records]]></article-title>
<source><![CDATA[ZooKeys]]></source>
<year>2017</year>
<volume>652</volume>
<page-range>129-79</page-range></nlm-citation>
</ref>
<ref id="B18">
<nlm-citation citation-type="book">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Cubero-Pardo]]></surname>
<given-names><![CDATA[P.]]></given-names>
</name>
</person-group>
<source><![CDATA[Distribución y patrones de actividad del bufeo (Tursiops truncatus) y el delfín manchado (Stenella attenuata) en el Golfo Dulce, en relación con variables ambientales]]></source>
<year>1998</year>
<publisher-loc><![CDATA[San José, Costa Rica ]]></publisher-loc>
<publisher-name><![CDATA[Universidad de Costa Rica]]></publisher-name>
</nlm-citation>
</ref>
<ref id="B19">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Fiedler]]></surname>
<given-names><![CDATA[P. C.]]></given-names>
</name>
<name>
<surname><![CDATA[Philbrick]]></surname>
<given-names><![CDATA[V.]]></given-names>
</name>
<name>
<surname><![CDATA[Chavez]]></surname>
<given-names><![CDATA[F. P.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Oceanic upwelling and productivity in the Eastern Tropical Pacific]]></article-title>
<source><![CDATA[Limnology and Oceanography]]></source>
<year>1991</year>
<volume>36</volume>
<numero>8</numero>
<issue>8</issue>
<page-range>1834-50</page-range></nlm-citation>
</ref>
<ref id="B20">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Fonseca]]></surname>
<given-names><![CDATA[L. G.]]></given-names>
</name>
<name>
<surname><![CDATA[Murillo]]></surname>
<given-names><![CDATA[G. A.]]></given-names>
</name>
<name>
<surname><![CDATA[Guadamúz]]></surname>
<given-names><![CDATA[L.]]></given-names>
</name>
<name>
<surname><![CDATA[Spínola]]></surname>
<given-names><![CDATA[R. M.]]></given-names>
</name>
<name>
<surname><![CDATA[Valverde]]></surname>
<given-names><![CDATA[R. A.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Downward but stable trend in the abundance of arribada olive ridley sea turtles (Lepidochelys olivacea) at Nancite Beach, Costa Rica (1971-2007)]]></article-title>
<source><![CDATA[Chelonian Conservation and Biology]]></source>
<year>2009</year>
<volume>8</volume>
<numero>1</numero>
<issue>1</issue>
<page-range>19-27</page-range></nlm-citation>
</ref>
<ref id="B21">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Fonseca]]></surname>
<given-names><![CDATA[L. G.]]></given-names>
</name>
<name>
<surname><![CDATA[Santidrián-Tomillo]]></surname>
<given-names><![CDATA[P.]]></given-names>
</name>
<name>
<surname><![CDATA[Villachica]]></surname>
<given-names><![CDATA[W. N.]]></given-names>
</name>
<name>
<surname><![CDATA[Quirós]]></surname>
<given-names><![CDATA[W. M.]]></given-names>
</name>
<name>
<surname><![CDATA[Pesquero]]></surname>
<given-names><![CDATA[M.]]></given-names>
</name>
<name>
<surname><![CDATA[Heidemeyer]]></surname>
<given-names><![CDATA[M.]]></given-names>
</name>
<name>
<surname><![CDATA[Valverde]]></surname>
<given-names><![CDATA[R. A.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Discovery of a major East Pacific green turtle (Chelonia mydas) nesting population in Northwest Costa Rica]]></article-title>
<source><![CDATA[Chelonian Conservation and Biology]]></source>
<year>2019</year>
<volume>17</volume>
<numero>2</numero>
<issue>2</issue>
<page-range>169</page-range></nlm-citation>
</ref>
<ref id="B22">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Gubbins]]></surname>
<given-names><![CDATA[C.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Use of home ranges by resident bottlenose dolphins (Tursiops truncatus) in a South Carolina estuary]]></article-title>
<source><![CDATA[Journal of Mammalogy]]></source>
<year>2002</year>
<volume>83</volume>
<numero>1</numero>
<issue>1</issue>
<page-range>178-87</page-range></nlm-citation>
</ref>
<ref id="B23">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Loría-Naranjo]]></surname>
<given-names><![CDATA[M.]]></given-names>
</name>
<name>
<surname><![CDATA[Samper-Villarreal]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
<name>
<surname><![CDATA[Cortés]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Structural complexity and species composition of Potrero Grande and Santa Elena mangrove forests in Santa Rosa National Park, North Pacific of Costa Rica]]></article-title>
<source><![CDATA[Revista de Biología Tropical]]></source>
<year>2014</year>
<volume>62</volume>
<numero>^s4</numero>
<issue>^s4</issue>
<supplement>4</supplement>
<page-range>S33-41</page-range></nlm-citation>
</ref>
<ref id="B24">
<nlm-citation citation-type="book">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Mann]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Unraveling the dynamics of social life: long-term studies and observation methods]]></article-title>
<person-group person-group-type="editor">
<name>
<surname><![CDATA[Mann]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
</person-group>
<source><![CDATA[Cetacean societies: field studies of dolphins and whales]]></source>
<year>2000</year>
<page-range>45-64</page-range><publisher-loc><![CDATA[Chicago ]]></publisher-loc>
<publisher-name><![CDATA[University of Chicago Press]]></publisher-name>
</nlm-citation>
</ref>
<ref id="B25">
<nlm-citation citation-type="book">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Martin]]></surname>
<given-names><![CDATA[P.]]></given-names>
</name>
<name>
<surname><![CDATA[Bateson]]></surname>
<given-names><![CDATA[P.]]></given-names>
</name>
</person-group>
<source><![CDATA[Measuring behaviour: an introdution guide]]></source>
<year>2007</year>
<edition>3</edition>
<publisher-loc><![CDATA[Cambridge ]]></publisher-loc>
<publisher-name><![CDATA[Cambridge University Press]]></publisher-name>
</nlm-citation>
</ref>
<ref id="B26">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Martínez-Fernández]]></surname>
<given-names><![CDATA[D.]]></given-names>
</name>
<name>
<surname><![CDATA[Montero-Cordero]]></surname>
<given-names><![CDATA[A.]]></given-names>
</name>
<name>
<surname><![CDATA[May-Collado]]></surname>
<given-names><![CDATA[L.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Cetáceos de las aguas costeras del Pacífico Norte y Sur de Costa Rica]]></article-title>
<source><![CDATA[Revista de Biología Tropical]]></source>
<year>2011</year>
<volume>59</volume>
<numero>1</numero>
<issue>1</issue>
<page-range>283-90</page-range></nlm-citation>
</ref>
<ref id="B27">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Martínez-Fernández]]></surname>
<given-names><![CDATA[D.]]></given-names>
</name>
<name>
<surname><![CDATA[Montero-Cordero]]></surname>
<given-names><![CDATA[A.]]></given-names>
</name>
<name>
<surname><![CDATA[Palacios-Alfaro]]></surname>
<given-names><![CDATA[D.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Áreas de congregación de cetáceos en el Pacífico Norte de Costa Rica: recomendaciones para la gestión del recurso]]></article-title>
<source><![CDATA[Revista de Biología Tropical]]></source>
<year>2014</year>
<volume>62</volume>
<numero>^s4</numero>
<issue>^s4</issue>
<supplement>4</supplement>
<page-range>S99-S108</page-range></nlm-citation>
</ref>
<ref id="B28">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[May-Collado]]></surname>
<given-names><![CDATA[L.]]></given-names>
</name>
<name>
<surname><![CDATA[Forcada]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Small-scale estimation of relative abundance for the coastal spotted dolphins (Stenella attenuata) in Costa Rica: the effect of habitat and seasonality]]></article-title>
<source><![CDATA[Revista de Biología Tropical]]></source>
<year>2012</year>
<volume>60</volume>
<numero>^s2</numero>
<issue>^s2</issue>
<supplement>2</supplement>
<page-range>S133-42</page-range></nlm-citation>
</ref>
<ref id="B29">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[May-Collado]]></surname>
<given-names><![CDATA[L.]]></given-names>
</name>
<name>
<surname><![CDATA[Gerrodette]]></surname>
<given-names><![CDATA[T.]]></given-names>
</name>
<name>
<surname><![CDATA[Calambokidis]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
<name>
<surname><![CDATA[Rasmussen]]></surname>
<given-names><![CDATA[K.]]></given-names>
</name>
<name>
<surname><![CDATA[Sereg]]></surname>
<given-names><![CDATA[I.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Patterns of cetacean sighting distribution in the Pacific Exclusive Economic Zone of Costa Rica based on data collected from 1979-2001]]></article-title>
<source><![CDATA[Revista de Biología Tropical]]></source>
<year>2005</year>
<volume>53</volume>
<numero>1-2</numero>
<issue>1-2</issue>
<page-range>249-63</page-range></nlm-citation>
</ref>
<ref id="B30">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[May-Collado]]></surname>
<given-names><![CDATA[L.]]></given-names>
</name>
<name>
<surname><![CDATA[Morales-Ramírez]]></surname>
<given-names><![CDATA[A.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Occurrence and behavioral patterns of the spotted coastal dolphin Stenella attenuata (Cetacea: Delphinidae) in the Gulf of Papagayo, Costa Rica]]></article-title>
<source><![CDATA[Revista de Biología Tropical]]></source>
<year>2005</year>
<volume>53</volume>
<numero>1-2</numero>
<issue>1-2</issue>
<page-range>265-76</page-range></nlm-citation>
</ref>
<ref id="B31">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Palacios-Alfaro]]></surname>
<given-names><![CDATA[J. D.]]></given-names>
</name>
<name>
<surname><![CDATA[Martínez-Fernández]]></surname>
<given-names><![CDATA[D.]]></given-names>
</name>
<name>
<surname><![CDATA[Sánchez]]></surname>
<given-names><![CDATA[C.]]></given-names>
</name>
<name>
<surname><![CDATA[Venegas-Li]]></surname>
<given-names><![CDATA[R.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Distribution and behavior of humpback whale (Megaptera novaeangliae Borowski, 1781) (Breeding Stock G), in Southern Pacific of Costa Rica]]></article-title>
<source><![CDATA[Journal of Cetacean Research Management]]></source>
<year>2009</year>
<volume>12</volume>
<numero>1</numero>
<issue>1</issue>
<page-range>1-8</page-range></nlm-citation>
</ref>
<ref id="B32">
<nlm-citation citation-type="book">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Rasmussen]]></surname>
<given-names><![CDATA[K.]]></given-names>
</name>
<name>
<surname><![CDATA[Calambokidis]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
<name>
<surname><![CDATA[Steiger]]></surname>
<given-names><![CDATA[G. H.]]></given-names>
</name>
</person-group>
<source><![CDATA[Humpback whales and other marine mammals off Costa Rica and surrounding waters, 1996-2002]]></source>
<year>2002</year>
<publisher-loc><![CDATA[San Francisco, California ]]></publisher-loc>
<publisher-name><![CDATA[Oceanic Society Expeditions]]></publisher-name>
</nlm-citation>
</ref>
<ref id="B33">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Rasmussen]]></surname>
<given-names><![CDATA[K.]]></given-names>
</name>
<name>
<surname><![CDATA[Calambokidis]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
<name>
<surname><![CDATA[Steiger]]></surname>
<given-names><![CDATA[G. H.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Distribution and migratory destinations of Humpback Whales off the Pacific Coast of Central America during the boreal winters of 1996-2003]]></article-title>
<source><![CDATA[Marine Mammal Science]]></source>
<year>2012</year>
<volume>28</volume>
<numero>3</numero>
<issue>3</issue>
<page-range>E267-79</page-range></nlm-citation>
</ref>
<ref id="B34">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Rasmussen]]></surname>
<given-names><![CDATA[K.]]></given-names>
</name>
<name>
<surname><![CDATA[Palacios]]></surname>
<given-names><![CDATA[D. M.]]></given-names>
</name>
<name>
<surname><![CDATA[Calambokidis]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
<name>
<surname><![CDATA[Saborío]]></surname>
<given-names><![CDATA[M. T.]]></given-names>
</name>
<name>
<surname><![CDATA[Dalla Rosa]]></surname>
<given-names><![CDATA[L.]]></given-names>
</name>
<name>
<surname><![CDATA[Secchi]]></surname>
<given-names><![CDATA[E. R.]]></given-names>
</name>
<name>
<surname><![CDATA[Stone]]></surname>
<given-names><![CDATA[G. S.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Southern Hemisphere humpback whales wintering off Central America: insights from water temperature into the longest mammalian migration]]></article-title>
<source><![CDATA[Biology Letters]]></source>
<year>2007</year>
<volume>3</volume>
<numero>3</numero>
<issue>3</issue>
<page-range>302-5</page-range></nlm-citation>
</ref>
<ref id="B35">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Schipper]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
<name>
<surname><![CDATA[Chanson]]></surname>
<given-names><![CDATA[J. S.]]></given-names>
</name>
<name>
<surname><![CDATA[Chiozza]]></surname>
<given-names><![CDATA[F.]]></given-names>
</name>
<name>
<surname><![CDATA[Cox]]></surname>
<given-names><![CDATA[N. A.]]></given-names>
</name>
<name>
<surname><![CDATA[Hoffmann]]></surname>
<given-names><![CDATA[M.]]></given-names>
</name>
<name>
<surname><![CDATA[Katariya]]></surname>
<given-names><![CDATA[V.]]></given-names>
</name>
<name>
<surname><![CDATA[Young]]></surname>
<given-names><![CDATA[B. E.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[The status of the world&#8217;s land and marine mammals: diversity, threat, and knowledge]]></article-title>
<source><![CDATA[Science]]></source>
<year>2008</year>
<volume>322</volume>
<numero>5899</numero>
<issue>5899</issue>
<page-range>225-30</page-range></nlm-citation>
</ref>
<ref id="B36">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Scott]]></surname>
<given-names><![CDATA[M. D.]]></given-names>
</name>
<name>
<surname><![CDATA[Cattanach]]></surname>
<given-names><![CDATA[K. L.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Diel patterns in aggregations of pelagic dolphins and tunas in the Eastern Pacific]]></article-title>
<source><![CDATA[Marine Mammal Science]]></source>
<year>1998</year>
<volume>14</volume>
<numero>3</numero>
<issue>3</issue>
<page-range>401-22</page-range></nlm-citation>
</ref>
<ref id="B37">
<nlm-citation citation-type="book">
<collab>SINAC</collab>
<source><![CDATA[GRUAS II: Propuesta de ordenamiento territorial para la conservación de la biodiversidad de Costa Rica]]></source>
<year>2008</year>
<publisher-loc><![CDATA[San José, Costa Rica ]]></publisher-loc>
<publisher-name><![CDATA[Ministerio de Ambiente y Energía]]></publisher-name>
</nlm-citation>
</ref>
<ref id="B38">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Steiger]]></surname>
<given-names><![CDATA[G. H.]]></given-names>
</name>
<name>
<surname><![CDATA[Calambokidis]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
<name>
<surname><![CDATA[Sears]]></surname>
<given-names><![CDATA[R.]]></given-names>
</name>
<name>
<surname><![CDATA[Balcomb]]></surname>
<given-names><![CDATA[K. C.]]></given-names>
</name>
<name>
<surname><![CDATA[Cubbage]]></surname>
<given-names><![CDATA[J. C.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Movement of Humpback Whales between California and Costa Rica]]></article-title>
<source><![CDATA[Marine Mammal Science]]></source>
<year>1991</year>
<volume>7</volume>
<numero>3</numero>
<issue>3</issue>
<page-range>306-10</page-range></nlm-citation>
</ref>
<ref id="B39">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Waylen]]></surname>
<given-names><![CDATA[P. R.]]></given-names>
</name>
<name>
<surname><![CDATA[Caviedes]]></surname>
<given-names><![CDATA[C. N.]]></given-names>
</name>
<name>
<surname><![CDATA[Quesada]]></surname>
<given-names><![CDATA[M. E.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Interannual variability of monthly precipitation in Costa Rica]]></article-title>
<source><![CDATA[Journal of Climate]]></source>
<year>1996</year>
<volume>9</volume>
<numero>10</numero>
<issue>10</issue>
<page-range>2606-13</page-range></nlm-citation>
</ref>
<ref id="B40">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Zamora-Trejos]]></surname>
<given-names><![CDATA[P.]]></given-names>
</name>
<name>
<surname><![CDATA[Cortés]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Los manglares de Costa Rica: el Pacífico Norte]]></article-title>
<source><![CDATA[Revista de Biología Tropical]]></source>
<year>2009</year>
<volume>57</volume>
<numero>3</numero>
<issue>3</issue>
<page-range>473-88</page-range></nlm-citation>
</ref>
</ref-list>
</back>
</article>
