<?xml version="1.0" encoding="ISO-8859-1"?><article xmlns:mml="http://www.w3.org/1998/Math/MathML" xmlns:xlink="http://www.w3.org/1999/xlink" xmlns:xsi="http://www.w3.org/2001/XMLSchema-instance">
<front>
<journal-meta>
<journal-id>1659-4266</journal-id>
<journal-title><![CDATA[Cuadernos de Investigación UNED]]></journal-title>
<abbrev-journal-title><![CDATA[Cuadernos de Investigación UNED]]></abbrev-journal-title>
<issn>1659-4266</issn>
<publisher>
<publisher-name><![CDATA[Universidad Estatal a Distancia de Costa Rica]]></publisher-name>
</publisher>
</journal-meta>
<article-meta>
<article-id>S1659-42662022000200044</article-id>
<article-id pub-id-type="doi">10.22458/urj.v14i2.4029</article-id>
<title-group>
<article-title xml:lang="es"><![CDATA[Claves para la identificación de los peces de las aguas continentales e insulares de Costa Rica. Parte II: Cichliformes, Cichlidae (Mojarras, Guapotes, Tilapias y afines)]]></article-title>
<article-title xml:lang="en"><![CDATA[Identification keys for the fish of continental and insular waters of Costa Rica. Part II: Cichliformes, Cichlidae (Mojarras, Guapotes, Tilapias and allies)]]></article-title>
</title-group>
<contrib-group>
<contrib contrib-type="author">
<name>
<surname><![CDATA[Angulo]]></surname>
<given-names><![CDATA[Arturo]]></given-names>
</name>
<xref ref-type="aff" rid="Aff"/>
<xref ref-type="aff" rid="Aaf"/>
<xref ref-type="aff" rid="A a"/>
<xref ref-type="aff" rid="A3"/>
</contrib>
<contrib contrib-type="author">
<name>
<surname><![CDATA[Gil-León]]></surname>
<given-names><![CDATA[Jorge San]]></given-names>
</name>
<xref ref-type="aff" rid="Aff"/>
<xref ref-type="aff" rid="Aaf"/>
</contrib>
</contrib-group>
<aff id="Af1">
<institution><![CDATA[,Universidad de Costa Rica Escuela de Biología ]]></institution>
<addr-line><![CDATA[ ]]></addr-line>
<country>Costa Rica</country>
</aff>
<aff id="Af2">
<institution><![CDATA[,Universidad de Costa Rica Centro de Investigación en Biodiversidad y Ecología Tropical ]]></institution>
<addr-line><![CDATA[ ]]></addr-line>
<country>Costa Rica</country>
</aff>
<aff id="Af3">
<institution><![CDATA[,Universidad de Costa Rica Centro de Investigación en Ciencias del Mar y Limnología ]]></institution>
<addr-line><![CDATA[ ]]></addr-line>
<country>Costa Rica</country>
</aff>
<pub-date pub-type="pub">
<day>00</day>
<month>12</month>
<year>2022</year>
</pub-date>
<pub-date pub-type="epub">
<day>00</day>
<month>12</month>
<year>2022</year>
</pub-date>
<volume>14</volume>
<numero>2</numero>
<fpage>44</fpage>
<lpage>69</lpage>
<copyright-statement/>
<copyright-year/>
<self-uri xlink:href="http://www.scielo.sa.cr/scielo.php?script=sci_arttext&amp;pid=S1659-42662022000200044&amp;lng=en&amp;nrm=iso"></self-uri><self-uri xlink:href="http://www.scielo.sa.cr/scielo.php?script=sci_abstract&amp;pid=S1659-42662022000200044&amp;lng=en&amp;nrm=iso"></self-uri><self-uri xlink:href="http://www.scielo.sa.cr/scielo.php?script=sci_pdf&amp;pid=S1659-42662022000200044&amp;lng=en&amp;nrm=iso"></self-uri><abstract abstract-type="short" xml:lang="es"><p><![CDATA[Resumen Introducción: Los cíclidos (Cichlidae, Cichliformes) constituyen una de las familias de peces dulceacuícolas más diversas en el planeta. En Costa Rica cerca de un 10% del total de las especies conocidas pertenecen a este grupo. Objetivo: Proveer, a aquellas personas interesadas (i.e., estudiantes, académicos, investigadores, aficionados y otros), información actualizada y en un lenguaje amigable que actualice la composición, clasificación y nomenclatura, permitiendo además la correcta identificación, de la totalidad de especies de Cichlidae presentes en las aguas continentales de Costa Rica. Métodos: Se levantó un listado de especies considerando registros bibliográficos e información obtenida a través de trabajo de campo. A partir de esta lista se elaboró una clave dicotómica, con información complementaria sobre distribución, basada principalmente en caracteres anatómicos externos y de coloración. La información utilizada para la elaboración de la clave proviene de tres fuentes principales: literatura científica, revisión de material museográfico y trabajo de campo. Resultados: Se listaron y analizaron un total de 14 géneros y 28 especies. La clave resultante está compuesta por 27 pasos permitiendo la identificación de la totalidad de especies de cíclidos presentes en Costa Rica. Se incluye además un catálogo fotográfico y datos actualizados sobre la nomenclatura de las especies listadas. Conclusión: Con esta contribución se actualiza el estado de conocimiento sobre la taxonomía, nomenclatura, distribución regional y local de las especies de Cichlidae de Costa Rica. La clave permite a su vez la correcta identificación de la totalidad de las especies de la familia y servirá de base para futuros estudios a nivel local y regional.]]></p></abstract>
<abstract abstract-type="short" xml:lang="en"><p><![CDATA[Abstract Introduction: Cichlids (Cichlidae, Cichliformes) constitute one of the most diverse families of freshwater fishes on the planet. In Costa Rica about 10% of the total known species belong to this group. Objective: Provide, to those interested (i.e., students, academics, researchers, hobbysts and others), updated information (in a friendly language) that updates the composition, classification and nomenclature, as well as allowing the correct identification, of all species of the Cichlidae occurring in the continental waters of Costa Rica. Methods: A list of species was made considering bibliographic records and information obtained through field work. From this list, an dichotomous key was elaborated, including complementary information on distribution, based mainly on external anatomical characters and coloration patterns. The information used for the elaboration of the key came from three main sources: scientific literature, review of museum specimens and fieldwork. Results: A total of 14 genera and 28 species were listed and analyzed. The resulting key is composed of 27 steps allowing the identification of all species of Cichlidae of Costa Rica. This paper also includes a photographic catalog and updated data on the nomenclature of the species listed. Conclusion: With this contribution, the state of knowledge about the taxonomy, nomenclature, regional and local distribution of the Cichlidae of Costa Rica is updated. The key allows the correct identification of all species of the family and will serve as the basis for future local and regional studies.]]></p></abstract>
<kwd-group>
<kwd lng="en"><![CDATA[Neotropical fishes]]></kwd>
<kwd lng="en"><![CDATA[Middle America]]></kwd>
<kwd lng="en"><![CDATA[Morphology]]></kwd>
<kwd lng="en"><![CDATA[External anatomy]]></kwd>
<kwd lng="en"><![CDATA[Dichotomous key.]]></kwd>
<kwd lng="es"><![CDATA[Peces Neotropicales]]></kwd>
<kwd lng="es"><![CDATA[América Media]]></kwd>
<kwd lng="es"><![CDATA[Morfología]]></kwd>
<kwd lng="es"><![CDATA[Anatomía externa]]></kwd>
<kwd lng="es"><![CDATA[Clave dicotómica.]]></kwd>
</kwd-group>
</article-meta>
</front><back>
<ref-list>
<ref id="B1">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Angulo]]></surname>
<given-names><![CDATA[A.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Nombres comunes y técnicos de los peces de agua dulce de Costa Rica.]]></article-title>
<source><![CDATA[Revista de Filología y Lingüística de la Universidad de Costa Rica]]></source>
<year>2013</year>
<volume>39</volume>
<numero>2</numero>
<issue>2</issue>
<page-range>77-103</page-range></nlm-citation>
</ref>
<ref id="B2">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Angulo]]></surname>
<given-names><![CDATA[A.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[New records and range extensions to the Costa Rican freshwater fish fauna, with an updated checklist.]]></article-title>
<source><![CDATA[Zootaxa]]></source>
<year>2021</year>
<volume>5083</volume>
<numero>1</numero>
<issue>1</issue>
<page-range>1-72</page-range></nlm-citation>
</ref>
<ref id="B3">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Angulo]]></surname>
<given-names><![CDATA[A.]]></given-names>
</name>
<name>
<surname><![CDATA[Garita-Alvarado]]></surname>
<given-names><![CDATA[C. A.]]></given-names>
</name>
<name>
<surname><![CDATA[Bussing]]></surname>
<given-names><![CDATA[W.A.]]></given-names>
</name>
<name>
<surname><![CDATA[López]]></surname>
<given-names><![CDATA[M.I.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Annotated checklist of the freshwater fishes of continental and insular Costa Rica: additions and nomenclatural revisions.]]></article-title>
<source><![CDATA[Check List]]></source>
<year>2013</year>
<volume>9</volume>
<numero>5</numero>
<issue>5</issue>
<page-range>987-1019</page-range></nlm-citation>
</ref>
<ref id="B4">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Angulo]]></surname>
<given-names><![CDATA[A.]]></given-names>
</name>
<name>
<surname><![CDATA[Ramirez-Coghi]]></surname>
<given-names><![CDATA[A.R.]]></given-names>
</name>
<name>
<surname><![CDATA[López]]></surname>
<given-names><![CDATA[M.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Claves para la identificación de los peces de las aguas continentales e insulares de Costa Rica. Parte I: Familias.]]></article-title>
<source><![CDATA[Cuadernos de Investigación UNED]]></source>
<year>2021</year>
<volume>13</volume>
<numero>1</numero>
<issue>1</issue>
</nlm-citation>
</ref>
<ref id="B5">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Bussing]]></surname>
<given-names><![CDATA[W. A.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Taxonomy and biological aspects of the Central American cichlid fishes Cichlasoma sieboldii and C. tuba.]]></article-title>
<source><![CDATA[Revista de Biología Tropical]]></source>
<year>1975</year>
<volume>23</volume>
<numero>2</numero>
<issue>2</issue>
<page-range>189-211</page-range></nlm-citation>
</ref>
<ref id="B6">
<nlm-citation citation-type="book">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Bussing]]></surname>
<given-names><![CDATA[W. A.]]></given-names>
</name>
</person-group>
<source><![CDATA[Peces de las aguas continentales de Costa Rica/Freshwater fishes of Costa Rica.]]></source>
<year>1998</year>
<publisher-name><![CDATA[Editorial de la Universidad de Costa Rica]]></publisher-name>
</nlm-citation>
</ref>
<ref id="B7">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Chakrabarty]]></surname>
<given-names><![CDATA[P.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[A morphological phylogenetic analysis of Middle American cichlids with special emphasis on the section "Nandopsis" sensu Regan.]]></article-title>
<source><![CDATA[Miscellaneous Publications, Museum of Zoology, University of Michigan]]></source>
<year>2007</year>
<volume>198</volume>
<page-range>1-31</page-range></nlm-citation>
</ref>
<ref id="B8">
<nlm-citation citation-type="book">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Conkel]]></surname>
<given-names><![CDATA[D.]]></given-names>
</name>
</person-group>
<source><![CDATA[Cichlids of North &amp; Central America]]></source>
<year>1993</year>
<publisher-name><![CDATA[Tfh Pubns Inc]]></publisher-name>
</nlm-citation>
</ref>
<ref id="B9">
<nlm-citation citation-type="">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Fricke]]></surname>
<given-names><![CDATA[R.]]></given-names>
</name>
<name>
<surname><![CDATA[Eschmeyer]]></surname>
<given-names><![CDATA[W. N.]]></given-names>
</name>
<name>
<surname><![CDATA[van der Laan]]></surname>
<given-names><![CDATA[R.]]></given-names>
</name>
</person-group>
<source><![CDATA[Eschmeyer's Catalog of Fishes: Genera, Species, References.]]></source>
<year>2022</year>
</nlm-citation>
</ref>
<ref id="B10">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Kner]]></surname>
<given-names><![CDATA[R.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Eine Uebersicht der ichthyologischen Ausbeute des Herrn Professors Dr. Mor. Wagner in Central-Amerika.]]></article-title>
<source><![CDATA[Sitzungsberichte der Königlich Bayerischen Akademie der Wissenschaften zu München,]]></source>
<year>1863</year>
<volume>1863</volume>
<numero>2</numero>
<issue>2</issue>
<page-range>220-30</page-range></nlm-citation>
</ref>
<ref id="B11">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[McMahan]]></surname>
<given-names><![CDATA[C. D.]]></given-names>
</name>
<name>
<surname><![CDATA[Matamoros]]></surname>
<given-names><![CDATA[W. A.]]></given-names>
</name>
<name>
<surname><![CDATA[Barraza]]></surname>
<given-names><![CDATA[E.]]></given-names>
</name>
<name>
<surname><![CDATA[Kutz]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
<name>
<surname><![CDATA[Chakrabarty]]></surname>
<given-names><![CDATA[P.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Taxonomic status of the Lago Coatepeque endemic convict cichlid Amatitlania coatepeque (Teleostei: Cichlidae).]]></article-title>
<source><![CDATA[Copeia]]></source>
<year>2014</year>
<volume>2014</volume>
<numero>4</numero>
<issue>4</issue>
<page-range>633-8</page-range></nlm-citation>
</ref>
<ref id="B12">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[McMahan]]></surname>
<given-names><![CDATA[C. D.]]></given-names>
</name>
<name>
<surname><![CDATA[Matamoros]]></surname>
<given-names><![CDATA[W. A.]]></given-names>
</name>
<name>
<surname><![CDATA[Piller]]></surname>
<given-names><![CDATA[K. R.]]></given-names>
</name>
<name>
<surname><![CDATA[Chakrabarty]]></surname>
<given-names><![CDATA[P.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Taxonomy and systematics of the herichthyins (Cichlidae: Tribe Heroini), with the description of eight new Middle American Genera]]></article-title>
<source><![CDATA[Zootaxa]]></source>
<year>2015</year>
<volume>3999</volume>
<numero>2</numero>
<issue>2</issue>
<page-range>211-34</page-range></nlm-citation>
</ref>
<ref id="B13">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Regan]]></surname>
<given-names><![CDATA[C. T.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[A collection of freshwater fishes made by Mr. C. F. underwood in Costa Rica.]]></article-title>
<source><![CDATA[Annals and Magazine of Natural History]]></source>
<year>1908</year>
<volume>8</volume>
<numero>2</numero>
<issue>2</issue>
<page-range>455-64</page-range></nlm-citation>
</ref>
<ref id="B14">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Regan]]></surname>
<given-names><![CDATA[C. T.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Pisces]]></article-title>
<source><![CDATA[Biologia Centrali-Americana]]></source>
<year>1906</year>
<volume>8</volume>
<page-range>1-203</page-range></nlm-citation>
</ref>
<ref id="B15">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[&#344;í&#269;an]]></surname>
<given-names><![CDATA[O.]]></given-names>
</name>
<name>
<surname><![CDATA[Piálek]]></surname>
<given-names><![CDATA[L.]]></given-names>
</name>
<name>
<surname><![CDATA[Dragová]]></surname>
<given-names><![CDATA[K.]]></given-names>
</name>
<name>
<surname><![CDATA[Novák]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Diversity and evolution of the Middle American cichlid fishes (Teleostei: Cichlidae) with revised classification.]]></article-title>
<source><![CDATA[Vertebrate Zoology]]></source>
<year>2016</year>
<volume>66</volume>
<numero>1</numero>
<issue>1</issue>
<page-range>3-102</page-range></nlm-citation>
</ref>
<ref id="B16">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Schmitter-Soto]]></surname>
<given-names><![CDATA[J. J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[A systematic revision of the genus Archocentrus (Perciformes: Cichlidae), with the description of two new genera and six new species.]]></article-title>
<source><![CDATA[Zootaxa]]></source>
<year>2007</year>
<volume>1603</volume>
<numero>1</numero>
<issue>1</issue>
<page-range>1-78</page-range></nlm-citation>
</ref>
</ref-list>
</back>
</article>
