1659-4266 1659-4266 S1659-42662019000300451 10.22458/urj.v11i3.2618 Costa Rica Costa Rica 00 12 2019 00 12 2019 11 3 451 454

2000 1977 1993 2016 2005 1976 1980 1985 1930 2011 2001 Panthera 2015 2013 2007 2009 1951