<?xml version="1.0" encoding="ISO-8859-1"?><article xmlns:mml="http://www.w3.org/1998/Math/MathML" xmlns:xlink="http://www.w3.org/1999/xlink" xmlns:xsi="http://www.w3.org/2001/XMLSchema-instance">
<front>
<journal-meta>
<journal-id>1659-4266</journal-id>
<journal-title><![CDATA[Cuadernos de Investigación UNED]]></journal-title>
<abbrev-journal-title><![CDATA[Cuadernos de Investigación UNED]]></abbrev-journal-title>
<issn>1659-4266</issn>
<publisher>
<publisher-name><![CDATA[Universidad Estatal a Distancia de Costa Rica]]></publisher-name>
</publisher>
</journal-meta>
<article-meta>
<article-id>S1659-42662015000100089</article-id>
<title-group>
<article-title xml:lang="en"><![CDATA[A new species of Hemibrycon (Characiformes: Characidae), a freshwater fish from the Sierra Nevada of Santa Marta, Caribbean drainage, Colombia]]></article-title>
<article-title xml:lang="es"><![CDATA[Una nueva especie de Hemibrycon (Characiformes: Characidae) de la Sierra Nevada de Santa Marta, Cuenca del Caribe, Norte de Colombia]]></article-title>
</title-group>
<contrib-group>
<contrib contrib-type="author">
<name>
<surname><![CDATA[Garcfa-Aizate]]></surname>
<given-names><![CDATA[Carlos A.]]></given-names>
</name>
<xref ref-type="aff" rid="Aff"/>
</contrib>
<contrib contrib-type="author">
<name>
<surname><![CDATA[Roman-Valencia]]></surname>
<given-names><![CDATA[Cesar]]></given-names>
</name>
<xref ref-type="aff" rid="Aff"/>
</contrib>
<contrib contrib-type="author">
<name>
<surname><![CDATA[Taphorn]]></surname>
<given-names><![CDATA[Donald C.]]></given-names>
</name>
<xref ref-type="aff" rid="Aff"/>
</contrib>
</contrib-group>
<aff id="Af1">
<institution><![CDATA[,Universidad del Atlántico  ]]></institution>
<addr-line><![CDATA[ Barranquilla]]></addr-line>
<country>Colombia</country>
</aff>
<aff id="Af2">
<institution><![CDATA[,Universidad del Quindío Laboratorio de Icti ]]></institution>
<addr-line><![CDATA[ Armenia]]></addr-line>
<country>Colombia</country>
</aff>
<pub-date pub-type="pub">
<day>00</day>
<month>06</month>
<year>2015</year>
</pub-date>
<pub-date pub-type="epub">
<day>00</day>
<month>06</month>
<year>2015</year>
</pub-date>
<volume>7</volume>
<numero>1</numero>
<fpage>89</fpage>
<lpage>98</lpage>
<copyright-statement/>
<copyright-year/>
<self-uri xlink:href="http://www.scielo.sa.cr/scielo.php?script=sci_arttext&amp;pid=S1659-42662015000100089&amp;lng=en&amp;nrm=iso"></self-uri><self-uri xlink:href="http://www.scielo.sa.cr/scielo.php?script=sci_abstract&amp;pid=S1659-42662015000100089&amp;lng=en&amp;nrm=iso"></self-uri><self-uri xlink:href="http://www.scielo.sa.cr/scielo.php?script=sci_pdf&amp;pid=S1659-42662015000100089&amp;lng=en&amp;nrm=iso"></self-uri><abstract abstract-type="short" xml:lang="en"><p><![CDATA[Abstract Hemibrycon is a group of freshwater characid fish species first recognized by Gunther (1864) as a subgenus of Tetragonopterus, and now included in the subfamily Stevardiinae. There are 21 spe cies recorded from the rivers of Colombia, but only one species, H. santamartae, has been reported previously in drainages of the Sierra Nevada de Santa Marta. Hemibrycon sierraensis n. sp. is described from the Río Gaira, Caribbean coastal drainage. It is distinguished from all congeners by having in life a bright red adipose fin (vs. hyaline or trans parent) that turns dark brown in specimens preserved in alcohol. It dif fers from Hemibrycon beni, H. boquiae, H. brevispini, H. colombianus, H. mikrostiktos, H. metae, H. microformaa, H. palomae and H. rafaelense in having a vertically elongate humeral spot that extends 3-4 longitudinal scale series below the lateral line. It differs from H. divisorensis, H. pau-tensis and H. santamartae by having the last dorsal-fin ray unbranched; and from H. pautensis by the number of scale series between the lat eral line and the pelvic-fin insertions. It differs from H. divisorensis in the number of unbranched anal-fin rays, and the number of teeth on the dentary and maxilla. It differs from H. santamartae in the number of su-praneurals, caudal peduncle depth, head length and orbital diameter.]]></p></abstract>
<abstract abstract-type="short" xml:lang="es"><p><![CDATA[ResumenHemibrycon es un grupo de especies de peces caracidos de agua dulce inicialmente reconocido por Gunther (1864) como un subgenera de Tetragonopterus, actualmente incluido en la subfamilia Stevardiinae y con 21 especies que se distribuyen en los rfos de Colombia, y solo una de ellas H. santamartae, se ha reportado previamente en los drenajes de la Sierra Nevada de Santa Marta. Describimos una nueva especie hallada en Río Gaira en la Sierra Nevada de Santa Marta, vertiente del Caribe Colombiano: Hemibrycon sierraen-sis. La nueva especie se distingue de sus congeneres por presentar, en vivo, la aleta adiposa roja brillante (vs. hialina o transparente) y en al cohol la aleta adiposa marron oscura. Tambien se distingue de H. beni, H. boquiae, H. brevispini, H. colombianus, H. mikrostiktos, H. metae, H. microformaa, H. palomae y H. rafaelense por presentar una mancha hu meral que alcanza 3-4 escamas por debajo del canal latero sensorial. H. sierraensis se separa de H. divisorensis, H. pautensis y H. santamartae por presentar el ultimo radio de la aleta dorsal simple, y se diferencia de H. pautensis por el numero de escamas entre la lfnea lateral y las aletas pelvicas. Difiere de H. divisorensis en el numero de radios simples de la aleta anal, el numero de dientes del maxilar y dentaRfo. H. sierraensis se separa de H. santamartae por el numero de supraneurales, por la profundidad y la longitud del pedunculo caudal, por la longitud de la cabeza y por el diametro del ojo.]]></p></abstract>
<kwd-group>
<kwd lng="en"><![CDATA[Biodiversity]]></kwd>
<kwd lng="en"><![CDATA[tropical fish]]></kwd>
<kwd lng="en"><![CDATA[new taxon]]></kwd>
<kwd lng="es"><![CDATA[Biodiversidad]]></kwd>
<kwd lng="es"><![CDATA[pez tropical]]></kwd>
<kwd lng="es"><![CDATA[nuevo taxon]]></kwd>
</kwd-group>
</article-meta>
</front><back>
<ref-list>
<ref id="B1">
<nlm-citation citation-type="book">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Arcila-Mesa]]></surname>
<given-names><![CDATA[D. K.]]></given-names>
</name>
</person-group>
<source><![CDATA[Análisis filogenético y biogeográfico de las especies de Hemibrycon (Characiformes, Characidae)]]></source>
<year>2008</year>
<publisher-loc><![CDATA[Armenia, Colombia ]]></publisher-loc>
<publisher-name><![CDATA[Universidad del Quindío, Programa de Biología]]></publisher-name>
</nlm-citation>
</ref>
<ref id="B2">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Bertaco]]></surname>
<given-names><![CDATA[V. A.,]]></given-names>
</name>
<name>
<surname><![CDATA[Malabarba]]></surname>
<given-names><![CDATA[L.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[A review of the Cis-Andean species of Hemibrycon Gunther (Teleostei: Characiformes: Characidae: Stevardiinae) with des cription of two new species]]></article-title>
<source><![CDATA[Neotropical Ichthyology]]></source>
<year>2010</year>
<volume>8</volume>
<page-range>737-70</page-range></nlm-citation>
</ref>
<ref id="B3">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Bertaco]]></surname>
<given-names><![CDATA[V. A.]]></given-names>
</name>
<name>
<surname><![CDATA[Malabarba]]></surname>
<given-names><![CDATA[L. R.]]></given-names>
</name>
<name>
<surname><![CDATA[Hidalgo]]></surname>
<given-names><![CDATA[M.,]]></given-names>
</name>
<name>
<surname><![CDATA[Ortega]]></surname>
<given-names><![CDATA[H.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[A new species of Hemibrycon (Teleostei: Characiformes: Characidae) from the río Ucayali drainage, Sierraensis del Divisor, Peru]]></article-title>
<source><![CDATA[Neotropical Ichthyology]]></source>
<year>2007</year>
<volume>5</volume>
<page-range>251-7</page-range></nlm-citation>
</ref>
<ref id="B4">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Eigenmann]]></surname>
<given-names><![CDATA[C. H.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[The American Characidae. Part 4]]></article-title>
<source><![CDATA[Memoirs of the Museum of Comparative Zoology at Harvard College]]></source>
<year>1927</year>
<volume>43</volume>
<page-range>401-16</page-range></nlm-citation>
</ref>
<ref id="B5">
<nlm-citation citation-type="">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Eschmeyer]]></surname>
<given-names><![CDATA[W. N.,]]></given-names>
</name>
<name>
<surname><![CDATA[Fricke]]></surname>
<given-names><![CDATA[R.]]></given-names>
</name>
</person-group>
<source><![CDATA[Catalog of Fishes electronic version]]></source>
<year>2013</year>
</nlm-citation>
</ref>
<ref id="B6">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Gery]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Notes on the ichthyology of Surinam and other Guianas. The distribution pattern of the genus Hemibrycon, with a description of a new species from Surinam and incursion in to ecotaxonomy]]></article-title>
<source><![CDATA[Bulletin Aquatic Biology]]></source>
<year>1962</year>
<volume>3</volume>
<page-range>65-80</page-range></nlm-citation>
</ref>
<ref id="B7">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Gunther]]></surname>
<given-names><![CDATA[A.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Catalogue of the fishes in the British Museum. Catalogue of the physostomi, containing the families Siluridae, Characinidae, Haplochitonidae, Sternoptychidae, Scopelidae, Stomiatidae in the collec tion of the British Museum]]></article-title>
<source><![CDATA[Catalogue fishes]]></source>
<year>1864</year>
<volume>5</volume>
<page-range>1-455</page-range></nlm-citation>
</ref>
<ref id="B8">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Javonillo]]></surname>
<given-names><![CDATA[R.]]></given-names>
</name>
<name>
<surname><![CDATA[Malabarba]]></surname>
<given-names><![CDATA[L. R.]]></given-names>
</name>
<name>
<surname><![CDATA[Weitzman]]></surname>
<given-names><![CDATA[S. H.,]]></given-names>
</name>
<name>
<surname><![CDATA[Burns]]></surname>
<given-names><![CDATA[J. R.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Relationships among major lineages of characid fishes (Teleostei: Ostariophysi: Characiformes), based on molecular sequence data]]></article-title>
<source><![CDATA[Molecular Phylogenetics and Evolution]]></source>
<year>2010</year>
<volume>54</volume>
<page-range>498-51</page-range></nlm-citation>
</ref>
<ref id="B9">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Mirande]]></surname>
<given-names><![CDATA[J. M.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Phylogeny of the family Characidae (Teleostei: Characiformes): from characters to taxo nomy]]></article-title>
<source><![CDATA[Neotropical Ichthyology]]></source>
<year>2010</year>
<volume>8</volume>
<page-range>385-568</page-range></nlm-citation>
</ref>
<ref id="B10">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Roman-Valencia]]></surname>
<given-names><![CDATA[C.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Redescripción de Hemibrycon boquiae (Pisces: Characidae), especie endémica de la Quebrada Boquia, cuenca Río Quindío, Alto Cauca, Colombia]]></article-title>
<source><![CDATA[Dahlia]]></source>
<year>2001</year>
<volume>4</volume>
<page-range>27-32</page-range></nlm-citation>
</ref>
<ref id="B11">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Roman-Valencia]]></surname>
<given-names><![CDATA[C.,]]></given-names>
</name>
<name>
<surname><![CDATA[Ruiz-C]]></surname>
<given-names><![CDATA[R. I.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Una nueva especie de pez del genero Hemibrycon (Characiformes: Characidae) del Alto Río Atrato, Noroccidente de Colombia]]></article-title>
<source><![CDATA[Caldasia]]></source>
<year>2007</year>
<volume>29</volume>
<page-range>121-31</page-range></nlm-citation>
</ref>
<ref id="B12">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Roman-Valencia]]></surname>
<given-names><![CDATA[C.,]]></given-names>
</name>
<name>
<surname><![CDATA[Arcila-Mesa.]]></surname>
<given-names><![CDATA[D. K.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Five new spe cies Hemibrycon (Characiformes: Characidae) from the Magdalena River Basin, Colombia]]></article-title>
<source><![CDATA[Revista de Biología Tropical]]></source>
<year>2010</year>
<volume>58</volume>
<page-range>339-56</page-range></nlm-citation>
</ref>
<ref id="B13">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Roman-Valencia]]></surname>
<given-names><![CDATA[C.]]></given-names>
</name>
<name>
<surname><![CDATA[Vanegas-Ríos]]></surname>
<given-names><![CDATA[J. A.,]]></given-names>
</name>
<name>
<surname><![CDATA[Garcfa]]></surname>
<given-names><![CDATA[M. D.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Análisis comparado de las especies del genero Bryconamericus (Teleostei: Characidae) en la cuenca de los ríos Cauca-Magdalena y Ranchería, Colombia]]></article-title>
<source><![CDATA[Revista Mexicana de Biodiversidad]]></source>
<year>2009</year>
<month>a</month>
<volume>80</volume>
<page-range>465-82</page-range></nlm-citation>
</ref>
<ref id="B14">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Roman-Valencia]]></surname>
<given-names><![CDATA[C.]]></given-names>
</name>
<name>
<surname><![CDATA[Ruiz-C.]]></surname>
<given-names><![CDATA[R.]]></given-names>
</name>
<name>
<surname><![CDATA[Garcfa-Alzate]]></surname>
<given-names><![CDATA[C.,]]></given-names>
</name>
<name>
<surname><![CDATA[Taphorn]]></surname>
<given-names><![CDATA[D. C.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Hemibrycon santamartae: a new species from the Rancheria River of Eastern Caribbean Colombia (Characiformes: Characidae)]]></article-title>
<source><![CDATA[Revista de investigaciones de la Universidad del Quindío]]></source>
<year>2009</year>
<month>b</month>
<volume>19</volume>
<page-range>144-50</page-range></nlm-citation>
</ref>
<ref id="B15">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Roman-Valencia]]></surname>
<given-names><![CDATA[C.]]></given-names>
</name>
<name>
<surname><![CDATA[Garcfa-Alzate]]></surname>
<given-names><![CDATA[C.]]></given-names>
</name>
<name>
<surname><![CDATA[Ruiz-C.]]></surname>
<given-names><![CDATA[R.,]]></given-names>
</name>
<name>
<surname><![CDATA[Taphorn]]></surname>
<given-names><![CDATA[D. C.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[New species of Hemibrycon (Teleostei, Characiformes, Characidae) from the Roble River, Alto Cauca, Colombia, with a key to species known from the Magdalena - Cauca River Basin]]></article-title>
<source><![CDATA[Vertebrate Zoology]]></source>
<year>2010</year>
<volume>60</volume>
<page-range>99-105</page-range></nlm-citation>
</ref>
<ref id="B16">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Roman-Valencia]]></surname>
<given-names><![CDATA[C.]]></given-names>
</name>
<name>
<surname><![CDATA[Ruiz-C.]]></surname>
<given-names><![CDATA[R.]]></given-names>
</name>
<name>
<surname><![CDATA[Taphorn]]></surname>
<given-names><![CDATA[D. C.]]></given-names>
</name>
<name>
<surname><![CDATA[Mancera-Rodriguez]]></surname>
<given-names><![CDATA[N. J.,]]></given-names>
</name>
<name>
<surname><![CDATA[Garcfa-Alzate]]></surname>
<given-names><![CDATA[C.A.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Three new species of Hemibrycon (Characiformes: Characidae) from the Magdalena River Basin, Colombia.]]></article-title>
<source><![CDATA[Revista de Biología Tropical]]></source>
<year>2013</year>
<volume>61</volume>
<page-range>1365-87</page-range></nlm-citation>
</ref>
<ref id="B17">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Ruiz-C.]]></surname>
<given-names><![CDATA[R. I.,]]></given-names>
</name>
<name>
<surname><![CDATA[Roman-Valencia]]></surname>
<given-names><![CDATA[C.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Osteología de Astyanax aurocaudatus Eigenmann, 1913 (Pisces: Characidae), con notas sobre la validez de Carlastyanax Gery, 1972]]></article-title>
<source><![CDATA[Animal Biodiversity and Conservation]]></source>
<year>2006</year>
<volume>29</volume>
<page-range>49-64</page-range></nlm-citation>
</ref>
<ref id="B18">
<nlm-citation citation-type="">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Perez]]></surname>
<given-names><![CDATA[Sabaj]]></given-names>
</name>
<name>
<surname><![CDATA[M.]]></surname>
<given-names><![CDATA[H.]]></given-names>
</name>
</person-group>
<source><![CDATA[Standard symbolic codes for institutional resource collections in herpetology and ichthy ology: an online reference]]></source>
<year>2010</year>
</nlm-citation>
</ref>
<ref id="B19">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Song]]></surname>
<given-names><![CDATA[J.,]]></given-names>
</name>
<name>
<surname><![CDATA[Parenti]]></surname>
<given-names><![CDATA[L. R.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Clearing and staining whole fish specimens for simultaneous demonstration of bone, cartilage and nerves]]></article-title>
<source><![CDATA[Copeia]]></source>
<year>1995</year>
<page-range>114-8</page-range></nlm-citation>
</ref>
<ref id="B20">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Taylor]]></surname>
<given-names><![CDATA[W. R.,]]></given-names>
</name>
<name>
<surname><![CDATA[van Dyke]]></surname>
<given-names><![CDATA[G. C.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Revised procedures for staining and clearing small fishes and other vertebrates for bone and cartilage study]]></article-title>
<source><![CDATA[Cybium]]></source>
<year>1985</year>
<volume>9</volume>
<page-range>107-19</page-range></nlm-citation>
</ref>
<ref id="B21">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Vari]]></surname>
<given-names><![CDATA[R.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[The Neotropical fish family Ctenoluciidae (Teleostei: Ostariophysi: Characiformes): supra and in-trafamilial phylogenetic relationships, with a revisionary study]]></article-title>
<source><![CDATA[Smithsonian Contributions to Zoology]]></source>
<year>1995</year>
<volume>564</volume>
<page-range>1-96</page-range></nlm-citation>
</ref>
<ref id="B22">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Weitzman]]></surname>
<given-names><![CDATA[S. H.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[The osteology of Brycon meeki, a gen eralized characid fish, with an osteological definition of the family]]></article-title>
<source><![CDATA[Stanford Ichthyology Bulletin]]></source>
<year>1962</year>
<volume>8</volume>
<page-range>1-77</page-range></nlm-citation>
</ref>
</ref-list>
</back>
</article>
