1409-3871 1409-3871 S1409-38712019000200125 10.15517/lank.v19i2.38786 Costa Rica Costa Rica Switzerland 00 08 2019 00 08 2019 19 2 125 193

1944 13 4 4 1979 2010 10 2-3 2-3 9-30 2012 2015 22 4 4 145-7 Field Museum of Natural History 2018 Harvard University 2018 a Harvard University 2018 b Harvard University 2018 c Kew Botanic Gardens 2019 1926 9 80 80 912-58 2006 6 3 3 91-4 Natural History Museum 2013 2009 9 1-2 1-2 1-268 1995 2013 47 5-145 1897 2002 3 1-146