<?xml version="1.0" encoding="ISO-8859-1"?><article xmlns:mml="http://www.w3.org/1998/Math/MathML" xmlns:xlink="http://www.w3.org/1999/xlink" xmlns:xsi="http://www.w3.org/2001/XMLSchema-instance">
<front>
<journal-meta>
<journal-id>0256-7024</journal-id>
<journal-title><![CDATA[Revista Geológica de América Central]]></journal-title>
<abbrev-journal-title><![CDATA[Rev. Geol. Amér. Central]]></abbrev-journal-title>
<issn>0256-7024</issn>
<publisher>
<publisher-name><![CDATA[Universidad de Costa Rica]]></publisher-name>
</publisher>
</journal-meta>
<article-meta>
<article-id>S0256-70242022000100067</article-id>
<article-id pub-id-type="doi">10.15517/rgac.v66i0.49763</article-id>
<title-group>
<article-title xml:lang="es"><![CDATA[Cormohipparion quinni (Mammalia, Equidae, Hipparionini) del Mioceno Medio (Barstoviano Tardío) del Norte de Costa Rica]]></article-title>
<article-title xml:lang="en"><![CDATA[Cormohipparion quinni (Mammalia, Equidae, Hipparionini) from the Middle Miocene (Late Barstovian) of Northern Costa Rica]]></article-title>
</title-group>
<contrib-group>
<contrib contrib-type="author">
<name>
<surname><![CDATA[Miller]]></surname>
<given-names><![CDATA[Stephanie]]></given-names>
</name>
<xref ref-type="aff" rid="Aff"/>
</contrib>
<contrib contrib-type="author">
<name>
<surname><![CDATA[Valerio]]></surname>
<given-names><![CDATA[Ana L.]]></given-names>
</name>
<xref ref-type="aff" rid="Aff"/>
</contrib>
<contrib contrib-type="author">
<name>
<surname><![CDATA[Sudasassi]]></surname>
<given-names><![CDATA[Arturo]]></given-names>
</name>
<xref ref-type="aff" rid="Aff"/>
</contrib>
<contrib contrib-type="author">
<name>
<surname><![CDATA[Laurito]]></surname>
<given-names><![CDATA[César A.]]></given-names>
</name>
<xref ref-type="aff" rid="Aff"/>
</contrib>
</contrib-group>
<aff id="Af1">
<institution><![CDATA[,Investigadores independientes  ]]></institution>
<addr-line><![CDATA[ ]]></addr-line>
<country>Costa Rica</country>
</aff>
<aff id="Af2">
<institution><![CDATA[,Museo Nacional de Costa Rica (MNCR) Departamento de Historia Natural Sección de Geología]]></institution>
<addr-line><![CDATA[ ]]></addr-line>
<country>Csoat Rica</country>
</aff>
<aff id="Af3">
<institution><![CDATA[,Museo Nacional de Costa Rica (MNCR) Departamento de Historia Natural ]]></institution>
<addr-line><![CDATA[ ]]></addr-line>
<country>Costa Rica</country>
</aff>
<pub-date pub-type="pub">
<day>00</day>
<month>06</month>
<year>2022</year>
</pub-date>
<pub-date pub-type="epub">
<day>00</day>
<month>06</month>
<year>2022</year>
</pub-date>
<numero>66</numero>
<fpage>67</fpage>
<lpage>75</lpage>
<copyright-statement/>
<copyright-year/>
<self-uri xlink:href="http://www.scielo.sa.cr/scielo.php?script=sci_arttext&amp;pid=S0256-70242022000100067&amp;lng=en&amp;nrm=iso"></self-uri><self-uri xlink:href="http://www.scielo.sa.cr/scielo.php?script=sci_abstract&amp;pid=S0256-70242022000100067&amp;lng=en&amp;nrm=iso"></self-uri><self-uri xlink:href="http://www.scielo.sa.cr/scielo.php?script=sci_pdf&amp;pid=S0256-70242022000100067&amp;lng=en&amp;nrm=iso"></self-uri><abstract abstract-type="short" xml:lang="es"><p><![CDATA[Resumen Por primera vez se registra la especie Cormohipparion quinniWoodburne 1996para el Barstoviano Tardío de América Central y Costa Rica. Este hallazgo aporta una nueva pista al patrón de distribución paleobiogeográfico de los mammíferos de América del Norte y su incursión en el Neotropico del sur de América Central durante el Mioceno Medio. El presente ejemplar representa el registro más austral de dicho taxón en el Mioceno Medio tardío en el Hemisferio Norte.]]></p></abstract>
<abstract abstract-type="short" xml:lang="en"><p><![CDATA[Abstract For the first time the species Cormohipparion quinniWoodburne 1996is recorded for the Late Barstovian of Central America and Costa Rica. This finding constitutes a new clue about the paleobiogeographical pattern of the Middle Miocene terrestrial mammals of North American and its incursion into the Neotropics of Southern Central America. This specimen represents the southernmost occurrence of this late Middle Miocene taxon at the North Hemisphere.]]></p></abstract>
<kwd-group>
<kwd lng="es"><![CDATA[hipparioninos]]></kwd>
<kwd lng="es"><![CDATA[Formación Venado]]></kwd>
<kwd lng="es"><![CDATA[Barstoviano Tardío]]></kwd>
<kwd lng="es"><![CDATA[Alajuela]]></kwd>
<kwd lng="es"><![CDATA[Costa Rica]]></kwd>
<kwd lng="en"><![CDATA[hipparionins]]></kwd>
<kwd lng="en"><![CDATA[Venado Formation]]></kwd>
<kwd lng="en"><![CDATA[Late Barstovian]]></kwd>
<kwd lng="en"><![CDATA[Alajuela]]></kwd>
<kwd lng="en"><![CDATA[Costa Rica]]></kwd>
</kwd-group>
</article-meta>
</front><back>
<ref-list>
<ref id="B1">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Bravo-Cuevas]]></surname>
<given-names><![CDATA[V. M.]]></given-names>
</name>
<name>
<surname><![CDATA[Ferrusquía-Villafranca]]></surname>
<given-names><![CDATA[I.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Merychippus (Mammalia, Perissodactyla, Equidae) from the Middle Miocene of state of Oaxaca, southeastern Mexico]]></article-title>
<source><![CDATA[Geobios]]></source>
<year>2006</year>
<volume>39</volume>
<numero>6</numero>
<issue>6</issue>
<page-range>771-84</page-range></nlm-citation>
</ref>
<ref id="B2">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Bravo-Cuevas]]></surname>
<given-names><![CDATA[V. M.]]></given-names>
</name>
<name>
<surname><![CDATA[Priego-Vargas]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Dietary evaluation of a hipparionin horse population from the middle Miocene of Oaxaca, southeastern Mexico]]></article-title>
<source><![CDATA[Revista Mexicana de Ciencias Geológicas]]></source>
<year>2009</year>
<volume>26</volume>
<numero>2</numero>
<issue>2</issue>
<page-range>356-66</page-range></nlm-citation>
</ref>
<ref id="B3">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Ferrusquía-Villlafranca]]></surname>
<given-names><![CDATA[I.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Mexico&#8217;s Middle Miocene mammalian assemblages: an overview]]></article-title>
<source><![CDATA[Bulletin of the American Museum of Natural History]]></source>
<year>2003</year>
<volume>13</volume>
<page-range>321-47</page-range></nlm-citation>
</ref>
<ref id="B4">
<nlm-citation citation-type="book">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Huapaya]]></surname>
<given-names><![CDATA[S.]]></given-names>
</name>
<name>
<surname><![CDATA[Rojas]]></surname>
<given-names><![CDATA[M.]]></given-names>
</name>
<name>
<surname><![CDATA[Rodríguez]]></surname>
<given-names><![CDATA[E.]]></given-names>
</name>
<name>
<surname><![CDATA[Jara]]></surname>
<given-names><![CDATA[D.]]></given-names>
</name>
<name>
<surname><![CDATA[Castro]]></surname>
<given-names><![CDATA[J. P.]]></given-names>
</name>
<name>
<surname><![CDATA[Segura]]></surname>
<given-names><![CDATA[G.]]></given-names>
</name>
<name>
<surname><![CDATA[Vázquez]]></surname>
<given-names><![CDATA[A.]]></given-names>
</name>
<name>
<surname><![CDATA[Pérez]]></surname>
<given-names><![CDATA[E. A.]]></given-names>
</name>
</person-group>
<source><![CDATA[Informe técnico financiero 2017]]></source>
<year>2017</year>
<publisher-loc><![CDATA[San José ]]></publisher-loc>
<publisher-name><![CDATA[Departamento de Investigación de la Dirección de Geología y Minas]]></publisher-name>
</nlm-citation>
</ref>
<ref id="B5">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Hulbert]]></surname>
<given-names><![CDATA[R. C.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Calippus and Protohippus (Mammalia, Perissodactyla, Equidae) from the Miocene (Barstovian-Early Hemphillian) of the Gulf Coastal Plain]]></article-title>
<source><![CDATA[Bulletin of the Florida State Museum Biological Sciences]]></source>
<year>1988</year>
<month>a</month>
<volume>32</volume>
<numero>3</numero>
<issue>3</issue>
<page-range>221-340</page-range></nlm-citation>
</ref>
<ref id="B6">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Hulbert]]></surname>
<given-names><![CDATA[R. C.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Cormohipparion and Hipparion (Mammalia, Perissodactyla, Equidae) from the Late Neogene of Florida]]></article-title>
<source><![CDATA[Bulletin of the Florida State Museum Biological Sciences]]></source>
<year>1988</year>
<month>b</month>
<volume>32</volume>
<numero>5</numero>
<issue>5</issue>
<page-range>229-338</page-range></nlm-citation>
</ref>
<ref id="B7">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Janis]]></surname>
<given-names><![CDATA[C. M.]]></given-names>
</name>
<name>
<surname><![CDATA[Damuth]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
<name>
<surname><![CDATA[Theodor]]></surname>
<given-names><![CDATA[J. M.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[The species richness of Miocene browsers and implications from habitat type and primary productivity in the North American grassland biome]]></article-title>
<source><![CDATA[Palaeogeography, Palaeoclimatology, Palaeoecology]]></source>
<year>2004</year>
<volume>207</volume>
<page-range>371-98</page-range></nlm-citation>
</ref>
<ref id="B8">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Leidy]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Rhinoceras and Hippotherium from Florida]]></article-title>
<source><![CDATA[Proceedings, Academy of Natural Science of Philadelphia]]></source>
<year>1885</year>
<volume>37</volume>
<page-range>32-3</page-range></nlm-citation>
</ref>
<ref id="B9">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Leidy]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Fossil Bones of Florida]]></article-title>
<source><![CDATA[Proceedings, Academy of Natural Science of Philadelphia]]></source>
<year>1887</year>
<volume>39</volume>
<page-range>309-10</page-range></nlm-citation>
</ref>
<ref id="B10">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[MacFadden]]></surname>
<given-names><![CDATA[B. J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Systematics and Phylogeny of Hipparion, Neohipparion, Nannipus, and Cormohipparion (Mammalia, Equidae) from the Miocene and Pliocene of the New World]]></article-title>
<source><![CDATA[Bulletin of the American Museum Natural History]]></source>
<year>1984</year>
<volume>179</volume>
<numero>1</numero>
<issue>1</issue>
<page-range>1-196</page-range></nlm-citation>
</ref>
<ref id="B11">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[MacFadden]]></surname>
<given-names><![CDATA[B. J.]]></given-names>
</name>
<name>
<surname><![CDATA[Jones]]></surname>
<given-names><![CDATA[D. S.]]></given-names>
</name>
<name>
<surname><![CDATA[Jud]]></surname>
<given-names><![CDATA[N. A.]]></given-names>
</name>
<name>
<surname><![CDATA[Moreno-Bernal]]></surname>
<given-names><![CDATA[J. W.]]></given-names>
</name>
<name>
<surname><![CDATA[Morgan]]></surname>
<given-names><![CDATA[G. S.]]></given-names>
</name>
<name>
<surname><![CDATA[Portell]]></surname>
<given-names><![CDATA[R. W.]]></given-names>
</name>
<name>
<surname><![CDATA[Perez]]></surname>
<given-names><![CDATA[V. J.]]></given-names>
</name>
<name>
<surname><![CDATA[Moran]]></surname>
<given-names><![CDATA[S. M.]]></given-names>
</name>
<name>
<surname><![CDATA[Wood]]></surname>
<given-names><![CDATA[A. R.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Integrated Chronology, Flora and Faunas, and Paleoecology of the Alajuela Formation, Late Miocene of Panama]]></article-title>
<source><![CDATA[PLoS ONE]]></source>
<year>2017</year>
<volume>12</volume>
<numero>1</numero>
<issue>1</issue>
</nlm-citation>
</ref>
<ref id="B12">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Obando]]></surname>
<given-names><![CDATA[L. G.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Estratigrafía de la Formación Venado y rocas sobreyacentes (Mioceno-Reciente) Provincia de Alajuela, Costa Rica]]></article-title>
<source><![CDATA[Revista Geológica de América Central]]></source>
<year>1986</year>
<volume>5</volume>
<page-range>73-104</page-range></nlm-citation>
</ref>
<ref id="B13">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Pagnac]]></surname>
<given-names><![CDATA[D.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Additions to the vertebrate faunal assemblage of the Middle Miocene Fort Randall Formation in the vicinity of South Bijou Hill, South Dakota, U.S.A]]></article-title>
<source><![CDATA[The Mountain Geologist]]></source>
<year>2012</year>
<volume>49</volume>
<numero>1</numero>
<issue>1</issue>
<page-range>19-34</page-range></nlm-citation>
</ref>
<ref id="B14">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Quinn]]></surname>
<given-names><![CDATA[J. H.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Miocene Equids of the Texas Gulf Coastal Plain. Bureau of the Economic Geology]]></article-title>
<source><![CDATA[University of Texas Publication]]></source>
<year>1955</year>
<volume>5516</volume>
<page-range>1-102</page-range></nlm-citation>
</ref>
<ref id="B15">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Retallack]]></surname>
<given-names><![CDATA[G. J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Cenozoic paleoclimate on land in North America]]></article-title>
<source><![CDATA[Journal of Geology]]></source>
<year>2007</year>
<volume>115</volume>
<page-range>271-94</page-range></nlm-citation>
</ref>
<ref id="B16">
<nlm-citation citation-type="book">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Rojas]]></surname>
<given-names><![CDATA[M.]]></given-names>
</name>
</person-group>
<source><![CDATA[Mapa geológico Hoja Monterrey (3247-I)]]></source>
<year>2019</year>
<publisher-loc><![CDATA[San José ]]></publisher-loc>
<publisher-name><![CDATA[Dirección de Geología, MINAE, República de Costa Rica]]></publisher-name>
</nlm-citation>
</ref>
<ref id="B17">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Skinner]]></surname>
<given-names><![CDATA[M. F.]]></given-names>
</name>
<name>
<surname><![CDATA[MacFadden]]></surname>
<given-names><![CDATA[B. J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Cormohipparion n. gen. (Mammalia, Equidae) from the North American Miocene (Barstovian-Clarendonian)]]></article-title>
<source><![CDATA[Journal of Paleontology]]></source>
<year>1977</year>
<volume>51</volume>
<page-range>912-26</page-range></nlm-citation>
</ref>
<ref id="B18">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Stirton]]></surname>
<given-names><![CDATA[R. A.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Development of characters in horse teeth and the dental nomenclature]]></article-title>
<source><![CDATA[Journal of Mammalogy]]></source>
<year>1941</year>
<volume>22</volume>
<numero>4</numero>
<issue>4</issue>
<page-range>434-46</page-range></nlm-citation>
</ref>
<ref id="B19">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Strömberg]]></surname>
<given-names><![CDATA[C. A. E.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[The origin and spread of grass-dominated ecosystems in the late Tertiary of North America: preliminary results concerning the evolution of hypsodonty]]></article-title>
<source><![CDATA[Palaeogeography, Palaeoclimatology, Palaeoecology]]></source>
<year>2002</year>
<volume>177</volume>
<page-range>59-75</page-range></nlm-citation>
</ref>
<ref id="B20">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Strömberg]]></surname>
<given-names><![CDATA[C. A. E.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Using phytolith assemblages to reconstruct the origin and spread of grass-dominated habitats in the great plains of North America during the late Eocene to early Miocene]]></article-title>
<source><![CDATA[Palaeogeography, Palaeoclimatology, Palaeoecology]]></source>
<year>2004</year>
<volume>207</volume>
<page-range>239-75</page-range></nlm-citation>
</ref>
<ref id="B21">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Webb]]></surname>
<given-names><![CDATA[S. D.]]></given-names>
</name>
<name>
<surname><![CDATA[Perrigo]]></surname>
<given-names><![CDATA[S. C.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Late Cenozoic vertebrates from Honduras and El Salvador]]></article-title>
<source><![CDATA[Journal of Vertebrate Paleontology]]></source>
<year>1984</year>
<volume>4</volume>
<page-range>237-54</page-range></nlm-citation>
</ref>
<ref id="B22">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Woodburne]]></surname>
<given-names><![CDATA[M.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Reappraisal of the Cormohipparion from the Valentine Formation, Nebraska]]></article-title>
<source><![CDATA[American Museum Novitates]]></source>
<year>1996</year>
<volume>3163</volume>
<page-range>1-56</page-range></nlm-citation>
</ref>
<ref id="B23">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Woodburne]]></surname>
<given-names><![CDATA[M.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Phyletic diversification of the Cormohipparion occidentale Complex (Mammalia, Perissodactyla, Equidae). Late Miocene, North America, and the origin of the Old Word Hippotherium Datum]]></article-title>
<source><![CDATA[Bulletin of the American Museum Natural History]]></source>
<year>2007</year>
<volume>306</volume>
<page-range>1-306</page-range></nlm-citation>
</ref>
</ref-list>
</back>
</article>
