<?xml version="1.0" encoding="ISO-8859-1"?><article xmlns:mml="http://www.w3.org/1998/Math/MathML" xmlns:xlink="http://www.w3.org/1999/xlink" xmlns:xsi="http://www.w3.org/2001/XMLSchema-instance">
<front>
<journal-meta>
<journal-id>0034-7744</journal-id>
<journal-title><![CDATA[Revista de Biología Tropical]]></journal-title>
<abbrev-journal-title><![CDATA[Rev. biol. trop]]></abbrev-journal-title>
<issn>0034-7744</issn>
<publisher>
<publisher-name><![CDATA[Universidad de Costa Rica]]></publisher-name>
</publisher>
</journal-meta>
<article-meta>
<article-id>S0034-77442020000100218</article-id>
<article-id pub-id-type="doi">10.15517/rbt.v68i1.37424</article-id>
<title-group>
<article-title xml:lang="en"><![CDATA[Anatomy and shell shape variability in a land snail Bostryx torallyi (Stylommatophora: Bulimulidae)]]></article-title>
<article-title xml:lang="es"><![CDATA[Anatomía y variabilidad de la forma de la concha en el caracol terrestre Bostryx torallyi (Stylommatophora: Bulimulidae)]]></article-title>
</title-group>
<contrib-group>
<contrib contrib-type="author">
<name>
<surname><![CDATA[Miranda]]></surname>
<given-names><![CDATA[María José]]></given-names>
</name>
<xref ref-type="aff" rid="Aff"/>
</contrib>
</contrib-group>
<aff id="Af1">
<institution><![CDATA[,Universidad Nacional de Tucumán Instituto de Biodiversidad Neotropical ]]></institution>
<addr-line><![CDATA[ ]]></addr-line>
<country>Argentina</country>
</aff>
<pub-date pub-type="pub">
<day>00</day>
<month>03</month>
<year>2020</year>
</pub-date>
<pub-date pub-type="epub">
<day>00</day>
<month>03</month>
<year>2020</year>
</pub-date>
<volume>68</volume>
<numero>1</numero>
<fpage>218</fpage>
<lpage>229</lpage>
<copyright-statement/>
<copyright-year/>
<self-uri xlink:href="http://www.scielo.sa.cr/scielo.php?script=sci_arttext&amp;pid=S0034-77442020000100218&amp;lng=en&amp;nrm=iso"></self-uri><self-uri xlink:href="http://www.scielo.sa.cr/scielo.php?script=sci_abstract&amp;pid=S0034-77442020000100218&amp;lng=en&amp;nrm=iso"></self-uri><self-uri xlink:href="http://www.scielo.sa.cr/scielo.php?script=sci_pdf&amp;pid=S0034-77442020000100218&amp;lng=en&amp;nrm=iso"></self-uri><abstract abstract-type="short" xml:lang="en"><p><![CDATA[Abstract  Introduction: The gastropod Bostryx torallyi shows high variability in shell shape and coloration. Subspecies of this organism have been described based on shell characters but, since they were slightly different, they were synonymized afterwards. Until now, shell variability has been analyzed only descriptively and its anatomy is still unknown.  Objective: In this study, I provide anatomical information of B. torallyi and apply a geometric morphometric analysis to evaluate the shell shape variation among specimens.  Methods: To accomplish this, type material and numerous lots were examined and dissected out. Additionally, relative warp analysis, based on 9 landmarks in ventral view of the shell, was performed using 80 specimens of 9 localities from Bolivia and Argentina.  Results: According to our results, geometric morphometrics is a suitable method to evaluate differences in shell shape among localities; for instance, distinctions in the shell were noticeable between gastropods of low and high altitudes. On the other hand, it was established that the coloration of this species is independent of large-scale factors since the examined specimens came from environments with similar conditions. Furthermore, the sculpture of the protoconch and anatomy of B. torallyi coincided with the other Argentinian species of the genus.  Conclusions: Therefore, I concluded that a geometric morphometric analysis of shell shape is a good complement to traditional qualitative description of the characteristics of the shell in this species.]]></p></abstract>
<abstract abstract-type="short" xml:lang="es"><p><![CDATA[Resumen  Introducción: El gasterópodo Bostryx torallyi exhibe una gran variabilidad en la forma y coloración de su concha. Subespecies de este organismo han sido descritas con base en los caracteres de la concha; pero, dadas las sutiles diferencias, fueron luego sinonimizadas. Hasta el momento, la variabilidad de la concha ha sido analizada solo descriptivamente y su anatomía es aún desconocida.  Objetivo: En este estudio, los objetivos fueron:proveer información anatómica de B. torallyi y usar un análisis de morfometría geométrica para evaluar la variación de la forma de la concha entre especímenes.  Métodos: Para ello, se examinó y disectó material tipo y numerosos lotes. Adicionalmente, se realizaron análisis del &#8220;relative warp&#8221;, basados en nueve &#8220;landmarks&#8221; en vista ventral de la concha, empleando 80 especímenes de nueve localidades de Bolivia y Argentina.  Resultados: De acuerdo con los resultados, la morfometría geométrica resultó útil para demostrar diferencias en forma de la concha entre localidades; por ejemplo, se detectarondiferencias evidentes entre localidades de alta y baja altitud. Se determinó que la variación en la coloración de esta especie es independiente de factores a gran escala, dado que los especímenes examinados proceden de ambientes con condiciones similares. Por otro lado, la escultura de la protoconcha y la anatomía de B. torallyi resultaron coincidentes con la de otras especies de este género recolectadas en Argentina.  Conclusiones: Se concluyó que el análisis de morfometría geométrica de la forma de la concha fue un buen complemento a la descripción cualitativa tradicional de caracteres de concha.]]></p></abstract>
<kwd-group>
<kwd lng="en"><![CDATA[altitude]]></kwd>
<kwd lng="en"><![CDATA[intraspecific variability]]></kwd>
<kwd lng="en"><![CDATA[geometric morphometric]]></kwd>
<kwd lng="en"><![CDATA[relative warp]]></kwd>
<kwd lng="en"><![CDATA[Argentina]]></kwd>
<kwd lng="en"><![CDATA[Bolivia]]></kwd>
<kwd lng="es"><![CDATA[altitud]]></kwd>
<kwd lng="es"><![CDATA[variabilidad intraspecífica]]></kwd>
<kwd lng="es"><![CDATA[morfometría geométrica]]></kwd>
<kwd lng="es"><![CDATA[deformación relativa]]></kwd>
<kwd lng="es"><![CDATA[Argentina]]></kwd>
<kwd lng="es"><![CDATA[Bolivia]]></kwd>
</kwd-group>
</article-meta>
</front><back>
<ref-list>
<ref id="B1">
<nlm-citation citation-type="book">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Backeljau]]></surname>
<given-names><![CDATA[T.]]></given-names>
</name>
<name>
<surname><![CDATA[Baur]]></surname>
<given-names><![CDATA[A.]]></given-names>
</name>
<name>
<surname><![CDATA[Baur]]></surname>
<given-names><![CDATA[B.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Population and Conservation Genetics]]></article-title>
<person-group person-group-type="editor">
<name>
<surname><![CDATA[Barker]]></surname>
<given-names><![CDATA[G. M.]]></given-names>
</name>
</person-group>
<source><![CDATA[The biology of terrestrial Molluscs]]></source>
<year>2001</year>
<page-range>383-412</page-range><publisher-loc><![CDATA[Wallingford, United Kingdom ]]></publisher-loc>
<publisher-name><![CDATA[CABI Publishing]]></publisher-name>
</nlm-citation>
</ref>
<ref id="B2">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Cameron]]></surname>
<given-names><![CDATA[R. A. D.]]></given-names>
</name>
<name>
<surname><![CDATA[Pokryszko]]></surname>
<given-names><![CDATA[B. M.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Variation in Cepaea populations over 42 years: climate fluctuations destroy a topographical relationship of morph-frequencies]]></article-title>
<source><![CDATA[Biological Journal of the Linnean Society]]></source>
<year>2008</year>
<volume>95</volume>
<page-range>53-61</page-range></nlm-citation>
</ref>
<ref id="B3">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Carvajal-Rodríguez]]></surname>
<given-names><![CDATA[A.]]></given-names>
</name>
<name>
<surname><![CDATA[Conde-Padin]]></surname>
<given-names><![CDATA[P.]]></given-names>
</name>
<name>
<surname><![CDATA[Rolan-Álvarez]]></surname>
<given-names><![CDATA[E.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Decomposing shell form into size and shape by geometric morphometric methods in two sympatric ecotypes of Littorina saxatilis]]></article-title>
<source><![CDATA[Journal of Molluscan Studies]]></source>
<year>2005</year>
<volume>71</volume>
<page-range>313-8</page-range></nlm-citation>
</ref>
<ref id="B4">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Chiba]]></surname>
<given-names><![CDATA[S.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Ecological and morphological patterns in communities of land snails of the genus Mandarina from the Bonin Islands]]></article-title>
<source><![CDATA[Journal of Evolution Biology]]></source>
<year>2004</year>
<volume>17</volume>
<page-range>131-43</page-range></nlm-citation>
</ref>
<ref id="B5">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Cuezzo]]></surname>
<given-names><![CDATA[M. G.]]></given-names>
</name>
<name>
<surname><![CDATA[Miranda]]></surname>
<given-names><![CDATA[M. J.]]></given-names>
</name>
<name>
<surname><![CDATA[Ovando]]></surname>
<given-names><![CDATA[X. M. C.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Species catalogue of Orthalicoidea in Argentina (Gastropoda: Stylommatophora)]]></article-title>
<source><![CDATA[Malacologia]]></source>
<year>2013</year>
<volume>56</volume>
<page-range>135-91</page-range></nlm-citation>
</ref>
<ref id="B6">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Dillon]]></surname>
<given-names><![CDATA[R. T.]]></given-names>
</name>
<name>
<surname><![CDATA[Jacquemin]]></surname>
<given-names><![CDATA[S. J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[The Heritability of Shell Morphometrics in the Freshwater Pulmonate Gastropod Physa]]></article-title>
<source><![CDATA[PLoS ONE]]></source>
<year>2015</year>
<volume>10</volume>
<numero>4</numero>
<issue>4</issue>
</nlm-citation>
</ref>
<ref id="B7">
<nlm-citation citation-type="">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Di Rienzo]]></surname>
<given-names><![CDATA[J. A.]]></given-names>
</name>
<name>
<surname><![CDATA[Casanoves]]></surname>
<given-names><![CDATA[F.]]></given-names>
</name>
<name>
<surname><![CDATA[Balzarini]]></surname>
<given-names><![CDATA[M. G.]]></given-names>
</name>
<name>
<surname><![CDATA[González]]></surname>
<given-names><![CDATA[L.]]></given-names>
</name>
<name>
<surname><![CDATA[Tablada]]></surname>
<given-names><![CDATA[M.]]></given-names>
</name>
<name>
<surname><![CDATA[Robledo]]></surname>
<given-names><![CDATA[C. W.]]></given-names>
</name>
</person-group>
<source><![CDATA[InfoStat]]></source>
<year>2016</year>
</nlm-citation>
</ref>
<ref id="B8">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Haase]]></surname>
<given-names><![CDATA[M.]]></given-names>
</name>
<name>
<surname><![CDATA[Esch]]></surname>
<given-names><![CDATA[S.]]></given-names>
</name>
<name>
<surname><![CDATA[Misof]]></surname>
<given-names><![CDATA[B.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Local adaptation, refugial isolation and secondary contact of alpine populations of the land snail Arianta arbustorum]]></article-title>
<source><![CDATA[Journal of Molluscan Studies]]></source>
<year>2013</year>
<volume>79</volume>
<page-range>241-8</page-range></nlm-citation>
</ref>
<ref id="B9">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Hylton-Scott]]></surname>
<given-names><![CDATA[M. I.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Moluscos del biotopo del Cerro Colorado (Salta]]></article-title>
<source><![CDATA[Acta Zoológica Lilloana]]></source>
<year>1948</year>
<volume>6</volume>
<page-range>229-39</page-range></nlm-citation>
</ref>
<ref id="B10">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Márquez]]></surname>
<given-names><![CDATA[F.]]></given-names>
</name>
<name>
<surname><![CDATA[van der Molen]]></surname>
<given-names><![CDATA[S.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Intraspecific shell-shape variation in the razor clam Ensis macha along the Patagonian coast]]></article-title>
<source><![CDATA[Journal of Molluscan Studies]]></source>
<year>2011</year>
<volume>77</volume>
<numero>2</numero>
<issue>2</issue>
<page-range>123-8</page-range></nlm-citation>
</ref>
<ref id="B11">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Miquel]]></surname>
<given-names><![CDATA[S. E.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Las especies del género Bostryx Troschel, 1847 en la República Argentina (2da. y última parte) (Gastropoda: Stylommatophora: Bulimulidae)]]></article-title>
<source><![CDATA[Archiv für Molluskenkunde]]></source>
<year>1995</year>
<volume>124</volume>
<page-range>119-27</page-range></nlm-citation>
</ref>
<ref id="B12">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Miranda]]></surname>
<given-names><![CDATA[M. J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Cladistic analysis of the Argentinian species of the genus Bostryx (Gastropoda, Stylommatophora) based on morphological evidence]]></article-title>
<source><![CDATA[Papeis Avulsos de Zoologia]]></source>
<year>2016</year>
<volume>56</volume>
<numero>10</numero>
<issue>10</issue>
<page-range>109-33</page-range></nlm-citation>
</ref>
<ref id="B13">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Miranda]]></surname>
<given-names><![CDATA[M. J.]]></given-names>
</name>
<name>
<surname><![CDATA[Cuezzo]]></surname>
<given-names><![CDATA[M. G.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Taxonomic revision of Bostryx stelzneri species complex, with description of a new species (Gastropoda: Orthalicoidea: Bulimulidae)]]></article-title>
<source><![CDATA[American Malacolological Bulletin]]></source>
<year>2014</year>
<volume>32</volume>
<numero>1</numero>
<issue>1</issue>
<page-range>74-93</page-range></nlm-citation>
</ref>
<ref id="B14">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Ozgo]]></surname>
<given-names><![CDATA[M.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Cepaea nemoralis (L.) in Southeastern Poland: Association of morpho frequencies with habitat]]></article-title>
<source><![CDATA[Journal of Molluscan Studies]]></source>
<year>2004</year>
<volume>71</volume>
<page-range>93-103</page-range></nlm-citation>
</ref>
<ref id="B15">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Parodiz]]></surname>
<given-names><![CDATA[J. J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Contribuciones al conocimiento de los moluscos terrestres Sudamericanos, 5]]></article-title>
<source><![CDATA[Comunicaciones Zoológicas del Museo Historia Natural de Montevideo]]></source>
<year>1947</year>
<volume>2</volume>
<page-range>1-32</page-range></nlm-citation>
</ref>
<ref id="B16">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Pfenninger]]></surname>
<given-names><![CDATA[M.]]></given-names>
</name>
<name>
<surname><![CDATA[Magnin]]></surname>
<given-names><![CDATA[F.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Phenotypic evolution and hidden speciation in Candidula unifasciatassp. (Helicellinae, Gastropoda) inferred by 16svariation and quantitative shell traits]]></article-title>
<source><![CDATA[Molecular Ecology]]></source>
<year>2001</year>
<volume>10</volume>
<page-range>2541-54</page-range></nlm-citation>
</ref>
<ref id="B17">
<nlm-citation citation-type="book">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Pilsbry]]></surname>
<given-names><![CDATA[H. A.]]></given-names>
</name>
</person-group>
<source><![CDATA[Manual of Conchology, Pulmonat]]></source>
<year>1901</year>
<publisher-loc><![CDATA[Philadelphia, United States ]]></publisher-loc>
<publisher-name><![CDATA[Palala Press]]></publisher-name>
</nlm-citation>
</ref>
<ref id="B18">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Ploeger]]></surname>
<given-names><![CDATA[S.]]></given-names>
</name>
<name>
<surname><![CDATA[Breure]]></surname>
<given-names><![CDATA[A. S. H.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[A rapid procedure for preparation of radulae for routine research with the scanning electron microscope]]></article-title>
<source><![CDATA[Basteria]]></source>
<year>1977</year>
<volume>41</volume>
<page-range>47-52</page-range></nlm-citation>
</ref>
<ref id="B19">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Quintana]]></surname>
<given-names><![CDATA[M.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Catálogo preliminar de la malacofauna del Paraguay]]></article-title>
<source><![CDATA[Revista Museo Argentino de Ciencias Naturales, Zoología]]></source>
<year>1982</year>
<volume>11</volume>
<page-range>61-158</page-range></nlm-citation>
</ref>
<ref id="B20">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Ramírez]]></surname>
<given-names><![CDATA[R.]]></given-names>
</name>
<name>
<surname><![CDATA[Paredes]]></surname>
<given-names><![CDATA[C.]]></given-names>
</name>
<name>
<surname><![CDATA[Arenas]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Moluscos del Perú]]></article-title>
<source><![CDATA[Revista de Biología Tropical]]></source>
<year>2003</year>
<volume>51</volume>
<page-range>225-84</page-range></nlm-citation>
</ref>
<ref id="B21">
<nlm-citation citation-type="">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Rohlf]]></surname>
<given-names><![CDATA[F. J.]]></given-names>
</name>
</person-group>
<source><![CDATA[TpsDig]]></source>
<year>2017</year>
</nlm-citation>
</ref>
<ref id="B22">
<nlm-citation citation-type="">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Rohlf]]></surname>
<given-names><![CDATA[F. J.]]></given-names>
</name>
</person-group>
<source><![CDATA[TpsRewl]]></source>
<year>2017</year>
</nlm-citation>
</ref>
<ref id="B23">
<nlm-citation citation-type="">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Rohlf]]></surname>
<given-names><![CDATA[F. J.]]></given-names>
</name>
</person-group>
<source><![CDATA[TpsUtil]]></source>
<year>2019</year>
</nlm-citation>
</ref>
<ref id="B24">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Signorelli]]></surname>
<given-names><![CDATA[J. H.]]></given-names>
</name>
<name>
<surname><![CDATA[Márquez]]></surname>
<given-names><![CDATA[F.]]></given-names>
</name>
<name>
<surname><![CDATA[Pastorino]]></surname>
<given-names><![CDATA[G.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Phenotypic variation of south-western Atlantic clam Mactra isabelleana (Bivalvia: Mactridae)]]></article-title>
<source><![CDATA[Journal of the Marine Biological Association of the United Kingdom]]></source>
<year>2012</year>
<volume>93</volume>
<numero>2</numero>
<issue>2</issue>
<page-range>1-7</page-range></nlm-citation>
</ref>
<ref id="B25">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Smith]]></surname>
<given-names><![CDATA[U. E.]]></given-names>
</name>
<name>
<surname><![CDATA[Hendricks]]></surname>
<given-names><![CDATA[J. R.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Geometric Morphometric Character Suites as Phylogenetic Data: Extracting Phylogenetic Signal from Gastropod Shells]]></article-title>
<source><![CDATA[Systematic Biology]]></source>
<year>2013</year>
<volume>62</volume>
<numero>3</numero>
<issue>3</issue>
<page-range>366-85</page-range></nlm-citation>
</ref>
<ref id="B26">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Surmacki]]></surname>
<given-names><![CDATA[A.]]></given-names>
</name>
<name>
<surname><![CDATA[Ozarowska-Nowicka]]></surname>
<given-names><![CDATA[A.]]></given-names>
</name>
<name>
<surname><![CDATA[Rosin]]></surname>
<given-names><![CDATA[Z. M.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Color polymorphism in a land snail Cepaea nemoralis (Pulmonata: Helicidae) as viewed by potential avian predators]]></article-title>
<source><![CDATA[Naturwissenschaften]]></source>
<year>2007</year>
<volume>100</volume>
<page-range>533-40</page-range></nlm-citation>
</ref>
<ref id="B27">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Zischka]]></surname>
<given-names><![CDATA[R.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Catálogo de las conchas y caracoles bolivianos]]></article-title>
<source><![CDATA[Folia Universitaria]]></source>
<year>1953</year>
<volume>10</volume>
<page-range>69-85</page-range></nlm-citation>
</ref>
</ref-list>
</back>
</article>
