<?xml version="1.0" encoding="ISO-8859-1"?><article xmlns:mml="http://www.w3.org/1998/Math/MathML" xmlns:xlink="http://www.w3.org/1999/xlink" xmlns:xsi="http://www.w3.org/2001/XMLSchema-instance">
<front>
<journal-meta>
<journal-id>0034-7744</journal-id>
<journal-title><![CDATA[Revista de Biología Tropical]]></journal-title>
<abbrev-journal-title><![CDATA[Rev. biol. trop]]></abbrev-journal-title>
<issn>0034-7744</issn>
<publisher>
<publisher-name><![CDATA[Universidad de Costa Rica]]></publisher-name>
</publisher>
</journal-meta>
<article-meta>
<article-id>S0034-77442019000200259</article-id>
<article-id pub-id-type="doi">10.15517/rbt.v67i2supl.37240</article-id>
<title-group>
<article-title xml:lang="en"><![CDATA[Shifts in the diversity of an amphibian community from a premontane forest of San Ramón, Costa Rica]]></article-title>
<article-title xml:lang="es"><![CDATA[Cambios en la diversidad de una comunidad de anfibios en un bosque premontano de San Ramón, Costa Rica]]></article-title>
</title-group>
<contrib-group>
<contrib contrib-type="author">
<name>
<surname><![CDATA[Acosta-Chaves]]></surname>
<given-names><![CDATA[Víctor J.]]></given-names>
</name>
<xref ref-type="aff" rid="Aff"/>
<xref ref-type="aff" rid="Aaf"/>
</contrib>
<contrib contrib-type="author">
<name>
<surname><![CDATA[Madrigal-Elizondo]]></surname>
<given-names><![CDATA[Víctor]]></given-names>
</name>
<xref ref-type="aff" rid="Aff"/>
</contrib>
<contrib contrib-type="author">
<name>
<surname><![CDATA[Chaves]]></surname>
<given-names><![CDATA[Gerardo]]></given-names>
</name>
<xref ref-type="aff" rid="Aff"/>
</contrib>
<contrib contrib-type="author">
<name>
<surname><![CDATA[Morera-Chacón]]></surname>
<given-names><![CDATA[Brayan]]></given-names>
</name>
<xref ref-type="aff" rid="Aff"/>
</contrib>
<contrib contrib-type="author">
<name>
<surname><![CDATA[García-Rodríguez]]></surname>
<given-names><![CDATA[Adrián]]></given-names>
</name>
<xref ref-type="aff" rid="Aff"/>
<xref ref-type="aff" rid="Aaf"/>
</contrib>
<contrib contrib-type="author">
<name>
<surname><![CDATA[Bolaños]]></surname>
<given-names><![CDATA[Federico]]></given-names>
</name>
<xref ref-type="aff" rid="Aff"/>
</contrib>
</contrib-group>
<aff id="Af1">
<institution><![CDATA[,Universidad de Costa Rica  ]]></institution>
<addr-line><![CDATA[Paraíso Cartago]]></addr-line>
<country>Costa Rica</country>
</aff>
<aff id="Af2">
<institution><![CDATA[,Red Mesoamericana y del Caribe para la Conservación de Anfibios y Reptiles  ]]></institution>
<addr-line><![CDATA[ ]]></addr-line>
</aff>
<aff id="Af3">
<institution><![CDATA[,Universidad de Costa Rica  ]]></institution>
<addr-line><![CDATA[San Pedro ]]></addr-line>
<country>Costa Rica</country>
</aff>
<aff id="Af4">
<institution><![CDATA[,Universidad de Costa Rica  ]]></institution>
<addr-line><![CDATA[San Pedro San José]]></addr-line>
<country>Costa Rica</country>
</aff>
<aff id="Af5">
<institution><![CDATA[,Universidad Nacional  ]]></institution>
<addr-line><![CDATA[ Heredia]]></addr-line>
<country>Costa Rica</country>
</aff>
<aff id="Af6">
<institution><![CDATA[,Universidad Nacional Autónoma de México Instituto de Biología Departamento de Zoología]]></institution>
<addr-line><![CDATA[ Mexico City]]></addr-line>
<country>Mexico</country>
</aff>
<aff id="Af6">
<institution><![CDATA[,aff6  ]]></institution>
<addr-line><![CDATA[ ]]></addr-line>
</aff>
<pub-date pub-type="pub">
<day>00</day>
<month>04</month>
<year>2019</year>
</pub-date>
<pub-date pub-type="epub">
<day>00</day>
<month>04</month>
<year>2019</year>
</pub-date>
<volume>67</volume>
<numero>2</numero>
<fpage>259</fpage>
<lpage>273</lpage>
<copyright-statement/>
<copyright-year/>
<self-uri xlink:href="http://www.scielo.sa.cr/scielo.php?script=sci_arttext&amp;pid=S0034-77442019000200259&amp;lng=en&amp;nrm=iso"></self-uri><self-uri xlink:href="http://www.scielo.sa.cr/scielo.php?script=sci_abstract&amp;pid=S0034-77442019000200259&amp;lng=en&amp;nrm=iso"></self-uri><self-uri xlink:href="http://www.scielo.sa.cr/scielo.php?script=sci_pdf&amp;pid=S0034-77442019000200259&amp;lng=en&amp;nrm=iso"></self-uri><abstract abstract-type="short" xml:lang="en"><p><![CDATA[Abstract Biological communities are experiencing rapid shifts of composition in Neotropical ecosystems due to several factors causing population declines. However, emerging evidence has provided insights on the adaptive potential of multiple species to respond to illnesses and environmental pressures. In Costa Rica, the decline of amphibian populations is a remarkable example of these changes. Here we provide evidence of variation in the amphibian richness of a premontane forest of San Ramón (Costa Rica) across a ~30 year period. We also quantified changes in the composition and abundance of the leaf-litter frog community occurring in the same premontane forest, by comparing diversity data with a difference of ~18 years. We evaluated the similarity of species richness from 1980s to 2010s based on several sources, and the dissimilarity of species diversity in the site comparing 28 standardized surveys from 1994-1995 and 2011-2012. We compared the relative abundance of some frogs that inhabit the leaf-litter layer between these same periods. Our results show that there is more similarity in amphibian richness between 1980s and 2010s (~ 52 %) than between 1980s and 1990s (~ 40 %). The richness of leaf-litter anurans was ~ 65 % similar between 1990s and 2010s. The diversity of leaf-litter anuran was clearly different between 1994-1995 and 2011-2012, and it was clustered among those periods. We determined that the amphibian community in this premontane forest drastically changed: many species have disappeared, or gradually declined through the decades (e.g. Pristimantis ridens, P. cruentus, Craugastor bransfordii) as in other well studied localities of Costa Rica, while some few species flourished after being almost absent from the site in the 1990s (e.g. Craugastor crassidigitus, Lithobates warszewistchii). Currently dominant species such as C. crassidigitus would be using developed resistance against Bd-fungus as an advantage (apparent competition) in the premontane forest where the disease is more virulent than in lowlands. Our analysis supports the hypothesis of individualized responses of anuran populations under distinct site and elevations. We suggest to continue monitoring the amphibian communities of premontane tropical forests to understand how this ecosystem gradually resist and adapts to this catastrophic time of biodiversity loss.]]></p></abstract>
<abstract abstract-type="short" xml:lang="es"><p><![CDATA[Resumen Las comunidades biológicas están experimentando rápidos cambios de composición en los ecosistemas neotropicales debido a varios factores que causan disminuciones de las poblaciones. Sin embargo, la evidencia emergente ha proporcionado información sobre el potencial de adaptación de múltiples especies para responder a enfermedades y presiones ambientales. En Costa Rica, el declive de las poblaciones de anfibios es un ejemplo notable de estos cambios. Aquí proporcionamos evidencia de variación en la riqueza de anfibios de un bosque premontano de San Ramón (Costa Rica) en un período de ~ 30 años. También cuantificamos los cambios en el ensamblaje, y las oscilaciones poblacionales, de la comunidad de ranas de hojarasca que ocurren en el mismo bosque premontano, tras comparar datos de diversidad con una diferencia de ~ 18 años. Evaluamos la similitud de la riqueza de especies desde la década de 1980 hasta la de 2010 basados en varias fuentes, así como la disimilitud de la diversidad de especies en el sitio comparando 28 muestreos estandarizados entre 1994-1995 y 2011-2012. Comparamos la abundancia relativa de algunas ranas que habitan la hojarasca entre 1994-1995 y 2011-2012. Nuestros resultados muestran que existe una similitud un poco mayor en la riqueza de anfibios entre los años 1980 y 2010 que entre los 1980 y 1990 (~ 40 %). La riqueza de los anuros de la hojarasca fue aproximadamente un 65 % similar entre la década de 1990 y 2010. La diversidad de anuros de la hojarasca fue claramente diferente entre 1994-1995 y 2011-2012, y se agrupó entre esos períodos. Determinamos que la comunidad de anfibios en este bosque premontano cambió drásticamente: muchas especies han desaparecido o han disminuido gradualmente a lo largo de las décadas como en otras localidades bien estudiadas de Costa Rica (por ejemplo, Pristimantis ridens, P. cruentus, Craugastor bransfordii), mientras algunas pocas especies florecieron tras casi desaparecer de nuestro sitio en la década de 1990 (por ejemplo, Craugastor crassidigitus, Lithobates warszewistchii). Actualmente, las especies dominantes como C. crassidigitus podrían tener resistencia contra el hongo quitridio, desarrollada como una ventaja (competición aparente) en el bosque premontano donde la enfermedad es más virulenta que en las tierras bajas. Nuestro análisis apoya la hipótesis de las respuestas individualizadas de las poblaciones de anuros en distintos sitios y elevaciones. Sugerimos continuar con el monitoreo de las comunidades de anfibios de los bosques neotropicales premontanos para comprender cómo este ecosistema resiste gradualmente y se adapta a este momento catastrófico de pérdida de biodiversidad.]]></p></abstract>
<kwd-group>
<kwd lng="en"><![CDATA[Adaptative potential]]></kwd>
<kwd lng="en"><![CDATA[Alberto Manuel Brenes Biological Reserve]]></kwd>
<kwd lng="en"><![CDATA[apparent competition]]></kwd>
<kwd lng="en"><![CDATA[Craugastor crassidigitus]]></kwd>
<kwd lng="en"><![CDATA[host resistance]]></kwd>
<kwd lng="en"><![CDATA[Lithobates warszewistchii]]></kwd>
<kwd lng="es"><![CDATA[Competición aparente]]></kwd>
<kwd lng="es"><![CDATA[Craugastor crassidigitus]]></kwd>
<kwd lng="es"><![CDATA[Lithobates warszewistchii]]></kwd>
<kwd lng="es"><![CDATA[Reserva Biológica Alberto Manuel Brenes]]></kwd>
<kwd lng="es"><![CDATA[resistencia del hospedero]]></kwd>
<kwd lng="es"><![CDATA[potencial adaptativo]]></kwd>
</kwd-group>
</article-meta>
</front><back>
<ref-list>
<ref id="B1">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Abarca-Alvarado]]></surname>
<given-names><![CDATA[J. G.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Cambios en la estructura de la comunidad de anuros (Amphibia: Anura) en el Cerro Chompipe, Costa Rica]]></article-title>
<source><![CDATA[Cuadernos de Investigación UNED]]></source>
<year>2012</year>
<volume>4</volume>
<numero>1</numero>
<issue>1</issue>
<page-range>9-15</page-range></nlm-citation>
</ref>
<ref id="B2">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Abarca]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
<name>
<surname><![CDATA[Chaves]]></surname>
<given-names><![CDATA[G.]]></given-names>
</name>
<name>
<surname><![CDATA[García-Rodríguez]]></surname>
<given-names><![CDATA[A.]]></given-names>
</name>
<name>
<surname><![CDATA[Vargas]]></surname>
<given-names><![CDATA[R.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Reconsidering extinction: Rediscovery of Incilius holdridgei (Anura: Bufonidae) in Costa Rica after 25 years]]></article-title>
<source><![CDATA[Herpetological Review]]></source>
<year>2009</year>
<volume>41</volume>
<page-range>150-2</page-range></nlm-citation>
</ref>
<ref id="B3">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Acosta-Chaves]]></surname>
<given-names><![CDATA[F.]]></given-names>
</name>
<name>
<surname><![CDATA[Spínola]]></surname>
<given-names><![CDATA[R. M.,]]></given-names>
</name>
<name>
<surname><![CDATA[Chaves]]></surname>
<given-names><![CDATA[G.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Density, biomass, and phenology of Craugastor underwoodi Boulenger (1896) from middle elevation forests in the Orosi Valley, Costa Rica]]></article-title>
<source><![CDATA[Mesoamerican Herpetology]]></source>
<year>2016</year>
<volume>3</volume>
<numero>4</numero>
<issue>4</issue>
<page-range>900-8</page-range></nlm-citation>
</ref>
<ref id="B4">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Acosta-Chaves]]></surname>
<given-names><![CDATA[V. J.]]></given-names>
</name>
<name>
<surname><![CDATA[Chaves]]></surname>
<given-names><![CDATA[G.]]></given-names>
</name>
<name>
<surname><![CDATA[Abarca]]></surname>
<given-names><![CDATA[J. G.]]></given-names>
</name>
<name>
<surname><![CDATA[García-Rodríguez]]></surname>
<given-names><![CDATA[A.]]></given-names>
</name>
<name>
<surname><![CDATA[Bolaños]]></surname>
<given-names><![CDATA[F.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[A checklist of the amphibians and reptiles of Río Macho Biological Station, Provincia de Cartago, Costa Rica]]></article-title>
<source><![CDATA[Check List]]></source>
<year>2015</year>
<volume>11</volume>
<numero>6</numero>
<issue>6</issue>
<page-range>1784</page-range></nlm-citation>
</ref>
<ref id="B5">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Acosta-Chaves]]></surname>
<given-names><![CDATA[V. J.]]></given-names>
</name>
<name>
<surname><![CDATA[Cossel Jr]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Smilisca phaeota (Cope, 1862). Colonization]]></article-title>
<source><![CDATA[Mesoamerican Herpetology]]></source>
<year>2016</year>
<volume>3</volume>
<numero>3</numero>
<issue>3</issue>
<page-range>713-5</page-range></nlm-citation>
</ref>
<ref id="B6">
<nlm-citation citation-type="book">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Alvarado]]></surname>
<given-names><![CDATA[G.]]></given-names>
</name>
</person-group>
<source><![CDATA[Batrachochytrium dendrobatidis en anuros de la Cordillera de Talamanca: estudio estructural, molecular y ecológico]]></source>
<year>2012</year>
<publisher-loc><![CDATA[Costa Rica ]]></publisher-loc>
<publisher-name><![CDATA[Universidad de Costa Rica]]></publisher-name>
</nlm-citation>
</ref>
<ref id="B7">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Bickford]]></surname>
<given-names><![CDATA[D.]]></given-names>
</name>
<name>
<surname><![CDATA[Howard]]></surname>
<given-names><![CDATA[S. D.]]></given-names>
</name>
<name>
<surname><![CDATA[Ng]]></surname>
<given-names><![CDATA[D. J. J.]]></given-names>
</name>
<name>
<surname><![CDATA[Sheridan]]></surname>
<given-names><![CDATA[J. A.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Impacts of climate change on the amphibians and reptiles of Southeast Asia]]></article-title>
<source><![CDATA[Biodiversity and Conservation]]></source>
<year>2010</year>
<volume>19</volume>
<numero>4</numero>
<issue>4</issue>
<page-range>1043-62</page-range></nlm-citation>
</ref>
<ref id="B8">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Bolaños]]></surname>
<given-names><![CDATA[F.]]></given-names>
</name>
<name>
<surname><![CDATA[Ehmcke]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Anfibios y reptiles de la Reserva Biológica San Ramón, Costa Rica]]></article-title>
<source><![CDATA[Revista Pensamiento Actual]]></source>
<year>1996</year>
<volume>2</volume>
<page-range>107-12</page-range></nlm-citation>
</ref>
<ref id="B9">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Brenes]]></surname>
<given-names><![CDATA[L.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Supervivencia, crecimiento y dispersión de rametes de Warszewiczia uxpanapensis (=Elaeagia uxpanapensis, Rubiaceae), Reserva Biológica Alberto Manuel Brenes, Costa Rica]]></article-title>
<source><![CDATA[Revista Pensamiento Actual]]></source>
<year>2007</year>
<volume>7</volume>
<page-range>113-5</page-range></nlm-citation>
</ref>
<ref id="B10">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Castro-Cruz]]></surname>
<given-names><![CDATA[A.]]></given-names>
</name>
<name>
<surname><![CDATA[García-Fernández]]></surname>
<given-names><![CDATA[F.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Report on Lithobates vibicarius (Cope, 1894) (Anura, Ranidae) in Parque Nacional del Agua Juan Castro Blanco, Alajuela, Costa Rica]]></article-title>
<source><![CDATA[Froglog]]></source>
<year>2012</year>
<volume>100</volume>
<page-range>69-70</page-range></nlm-citation>
</ref>
<ref id="B11">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Chaves]]></surname>
<given-names><![CDATA[G.]]></given-names>
</name>
<name>
<surname><![CDATA[Zumbado-Ulate]]></surname>
<given-names><![CDATA[H.]]></given-names>
</name>
<name>
<surname><![CDATA[García-Rodríguez]]></surname>
<given-names><![CDATA[A.]]></given-names>
</name>
<name>
<surname><![CDATA[Gómez]]></surname>
<given-names><![CDATA[E.]]></given-names>
</name>
<name>
<surname><![CDATA[Vredenburg]]></surname>
<given-names><![CDATA[V. T.]]></given-names>
</name>
<name>
<surname><![CDATA[Ryan]]></surname>
<given-names><![CDATA[M. J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Rediscovery of the critically endangered streamside frog, Craugastor taurus (Craugastoridae), in Costa Rica]]></article-title>
<source><![CDATA[Tropical Conservation Science]]></source>
<year>2014</year>
<volume>7</volume>
<page-range>628-38</page-range></nlm-citation>
</ref>
<ref id="B12">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Colwell]]></surname>
<given-names><![CDATA[R. K.]]></given-names>
</name>
<name>
<surname><![CDATA[Brehm]]></surname>
<given-names><![CDATA[G.]]></given-names>
</name>
<name>
<surname><![CDATA[Cardelús]]></surname>
<given-names><![CDATA[C. L.]]></given-names>
</name>
<name>
<surname><![CDATA[Gilman]]></surname>
<given-names><![CDATA[A. C.]]></given-names>
</name>
<name>
<surname><![CDATA[Longino]]></surname>
<given-names><![CDATA[J. T.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Global warming, elevational range shifts, and lowland biotic attrition in the wet tropics]]></article-title>
<source><![CDATA[Science]]></source>
<year>2008</year>
<volume>322</volume>
<numero>5899</numero>
<issue>5899</issue>
<page-range>258-61</page-range></nlm-citation>
</ref>
<ref id="B13">
<nlm-citation citation-type="book">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Crump]]></surname>
<given-names><![CDATA[M. L.]]></given-names>
</name>
<name>
<surname><![CDATA[Scott]]></surname>
<given-names><![CDATA[N. J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Visual encounter surveys]]></article-title>
<person-group person-group-type="editor">
<name>
<surname><![CDATA[Heyer]]></surname>
<given-names><![CDATA[W. R.]]></given-names>
</name>
</person-group>
<source><![CDATA[Measuring and Monitoring Biological Diversity. Standard Methods for Amphibians]]></source>
<year>1994</year>
<page-range>84-92</page-range><publisher-loc><![CDATA[Washington and London ]]></publisher-loc>
<publisher-name><![CDATA[Smithsonian Institution Press]]></publisher-name>
</nlm-citation>
</ref>
<ref id="B14">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[González-Maya]]></surname>
<given-names><![CDATA[J. F.]]></given-names>
</name>
<name>
<surname><![CDATA[Belant]]></surname>
<given-names><![CDATA[J. L.]]></given-names>
</name>
<name>
<surname><![CDATA[Wyatt]]></surname>
<given-names><![CDATA[S. A.]]></given-names>
</name>
<name>
<surname><![CDATA[Schipper]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
<name>
<surname><![CDATA[Cardenal]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
<name>
<surname><![CDATA[Corrales]]></surname>
<given-names><![CDATA[D.]]></given-names>
</name>
<name>
<surname><![CDATA[Fischer]]></surname>
<given-names><![CDATA[A.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Renewing hope: the rediscovery of Atelopus varius in Costa Rica]]></article-title>
<source><![CDATA[Amphibia-Reptilia]]></source>
<year>2013</year>
<volume>34</volume>
<numero>4</numero>
<issue>4</issue>
<page-range>573-8</page-range></nlm-citation>
</ref>
<ref id="B15">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Hammer]]></surname>
<given-names><![CDATA[Ø.]]></given-names>
</name>
<name>
<surname><![CDATA[Harper]]></surname>
<given-names><![CDATA[D. A. T.]]></given-names>
</name>
<name>
<surname><![CDATA[Ryan]]></surname>
<given-names><![CDATA[P. D.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Paleontological statistics software: package for education and data analysis]]></article-title>
<source><![CDATA[Palaeontologia Electronica]]></source>
<year>2001</year>
<volume>4</volume>
<numero>1</numero>
<issue>1</issue>
<page-range>1-9</page-range></nlm-citation>
</ref>
<ref id="B16">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Hertz]]></surname>
<given-names><![CDATA[A.]]></given-names>
</name>
<name>
<surname><![CDATA[Lotzkat]]></surname>
<given-names><![CDATA[S.]]></given-names>
</name>
<name>
<surname><![CDATA[Carrizo]]></surname>
<given-names><![CDATA[A.]]></given-names>
</name>
<name>
<surname><![CDATA[Ponce]]></surname>
<given-names><![CDATA[M.]]></given-names>
</name>
<name>
<surname><![CDATA[Köhler]]></surname>
<given-names><![CDATA[G.]]></given-names>
</name>
<name>
<surname><![CDATA[Streit]]></surname>
<given-names><![CDATA[B.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Field notes on findings of threatened amphibian species in the central mountain range of western Panama]]></article-title>
<source><![CDATA[Amphibian and Reptile Conservation]]></source>
<year>2012</year>
<volume>6</volume>
<numero>2</numero>
<issue>2</issue>
<page-range>9-30</page-range></nlm-citation>
</ref>
<ref id="B17">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Hilje]]></surname>
<given-names><![CDATA[B.]]></given-names>
</name>
<name>
<surname><![CDATA[Aide]]></surname>
<given-names><![CDATA[T. M.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Recovery of amphibian species richness and composition in a chronosequence of secondary forests, Northeastern Costa Rica]]></article-title>
<source><![CDATA[Biological Conservation]]></source>
<year>2012</year>
<volume>146</volume>
<page-range>170-6</page-range></nlm-citation>
</ref>
<ref id="B18">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Ihaka]]></surname>
<given-names><![CDATA[R.]]></given-names>
</name>
<name>
<surname><![CDATA[Gentleman]]></surname>
<given-names><![CDATA[R.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[R: a language for data analysis and graphics]]></article-title>
<source><![CDATA[Journal of Computational and Graphical Statistics]]></source>
<year>1996</year>
<volume>5</volume>
<page-range>299-314</page-range></nlm-citation>
</ref>
<ref id="B19">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Jiménez]]></surname>
<given-names><![CDATA[R.]]></given-names>
</name>
<name>
<surname><![CDATA[Alvarado]]></surname>
<given-names><![CDATA[G.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Craugastor escoces (Anura: Craugastoridae) reappears after 30 years: rediscovery of an &#8220;extinct&#8221; Neotropical frog]]></article-title>
<source><![CDATA[Amphibia-Reptilia]]></source>
<year>2017</year>
<volume>38</volume>
<page-range>257-9</page-range></nlm-citation>
</ref>
<ref id="B20">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Kubicki]]></surname>
<given-names><![CDATA[B.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Rediscovery of the critically endangered frog, Craugastor angelicus, in Costa Rica]]></article-title>
<source><![CDATA[Mesoamerican Herpetology]]></source>
<year>2016</year>
<volume>3</volume>
<page-range>1070-1</page-range></nlm-citation>
</ref>
<ref id="B21">
<nlm-citation citation-type="book">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Leenders]]></surname>
<given-names><![CDATA[T.]]></given-names>
</name>
</person-group>
<source><![CDATA[Amphibians of Costa Rica. A Field Guide]]></source>
<year>2016</year>
<publisher-loc><![CDATA[Nueva York ]]></publisher-loc>
<publisher-name><![CDATA[Zona Tropical Publications]]></publisher-name>
</nlm-citation>
</ref>
<ref id="B22">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Lips]]></surname>
<given-names><![CDATA[K. R.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Decline of a tropical amphibian fauna]]></article-title>
<source><![CDATA[Conservation. Biology]]></source>
<year>1998</year>
<volume>12</volume>
<page-range>106-17</page-range></nlm-citation>
</ref>
<ref id="B23">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Lips]]></surname>
<given-names><![CDATA[K. R.]]></given-names>
</name>
<name>
<surname><![CDATA[Brem]]></surname>
<given-names><![CDATA[F.]]></given-names>
</name>
<name>
<surname><![CDATA[Brenes]]></surname>
<given-names><![CDATA[R.]]></given-names>
</name>
<name>
<surname><![CDATA[Reeve]]></surname>
<given-names><![CDATA[J. D.]]></given-names>
</name>
<name>
<surname><![CDATA[Alford]]></surname>
<given-names><![CDATA[R. A.]]></given-names>
</name>
<name>
<surname><![CDATA[Voyles]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
<name>
<surname><![CDATA[&#8230; Collins]]></surname>
<given-names><![CDATA[J. P.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Emerging infectious disease and the loss of biodiversity in a Neotropical amphibian community]]></article-title>
<source><![CDATA[Proceedings of the National Academy of Sciences of the United States of America]]></source>
<year>2006</year>
<volume>103</volume>
<numero>9</numero>
<issue>9</issue>
<page-range>3165-70</page-range></nlm-citation>
</ref>
<ref id="B24">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Lister]]></surname>
<given-names><![CDATA[B. C.]]></given-names>
</name>
<name>
<surname><![CDATA[Garcia]]></surname>
<given-names><![CDATA[A.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Climate-driven declines in arthropod abundance restructure a rainforest food web]]></article-title>
<source><![CDATA[Proceedings of the National Academy of Sciences]]></source>
<year>2018</year>
<volume>115</volume>
<numero>44</numero>
<issue>44</issue>
<page-range>E10397-406</page-range></nlm-citation>
</ref>
<ref id="B25">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Mendelson III]]></surname>
<given-names><![CDATA[J. R.]]></given-names>
</name>
<name>
<surname><![CDATA[Brodie Jr]]></surname>
<given-names><![CDATA[E. D.]]></given-names>
</name>
<name>
<surname><![CDATA[Malone]]></surname>
<given-names><![CDATA[J. H.]]></given-names>
</name>
<name>
<surname><![CDATA[Acevedo]]></surname>
<given-names><![CDATA[M. E.]]></given-names>
</name>
<name>
<surname><![CDATA[Baker]]></surname>
<given-names><![CDATA[M. A.]]></given-names>
</name>
<name>
<surname><![CDATA[Smatresk]]></surname>
<given-names><![CDATA[N. J.]]></given-names>
</name>
<name>
<surname><![CDATA[Campbell]]></surname>
<given-names><![CDATA[J. A.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Factors associated with the catastrophic decline of a cloud forest frog fauna in Guatemala]]></article-title>
<source><![CDATA[Revista de Biología Tropical]]></source>
<year>2004</year>
<volume>52</volume>
<numero>4</numero>
<issue>4</issue>
<page-range>991-1000</page-range></nlm-citation>
</ref>
<ref id="B26">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Morera-Chacón]]></surname>
<given-names><![CDATA[B. H.]]></given-names>
</name>
<name>
<surname><![CDATA[Jiménez-Castro]]></surname>
<given-names><![CDATA[J. E.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Primer reporte de Agalychnis callidryas (Hylidae) en la Reserva Biológica Alberto Manuel Brenes]]></article-title>
<source><![CDATA[Revista Pensamiento Actual]]></source>
<year>2017</year>
<volume>17</volume>
<numero>1</numero>
<issue>1</issue>
<page-range>59-60</page-range></nlm-citation>
</ref>
<ref id="B27">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Morera-Chacón]]></surname>
<given-names><![CDATA[B. H.]]></given-names>
</name>
<name>
<surname><![CDATA[Sánchez-Porras]]></surname>
<given-names><![CDATA[R.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Anfibios y reptiles de la Reserva Biológica Alberto Manuel Brenes]]></article-title>
<source><![CDATA[Revista Pensamiento Actual]]></source>
<year>2015</year>
<volume>15</volume>
<numero>25</numero>
<issue>25</issue>
<page-range>39-59</page-range></nlm-citation>
</ref>
<ref id="B28">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Moritz]]></surname>
<given-names><![CDATA[C.]]></given-names>
</name>
<name>
<surname><![CDATA[Patton]]></surname>
<given-names><![CDATA[J. L.]]></given-names>
</name>
<name>
<surname><![CDATA[Conroy]]></surname>
<given-names><![CDATA[C. J.]]></given-names>
</name>
<name>
<surname><![CDATA[Parra]]></surname>
<given-names><![CDATA[J. L.]]></given-names>
</name>
<name>
<surname><![CDATA[White]]></surname>
<given-names><![CDATA[G. C.]]></given-names>
</name>
<name>
<surname><![CDATA[Beissinger]]></surname>
<given-names><![CDATA[S. R.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Impact of a century of climate change on small-mammal communities in Yosemite National Park, USA]]></article-title>
<source><![CDATA[Science]]></source>
<year>2008</year>
<volume>322</volume>
<numero>5899</numero>
<issue>5899</issue>
<page-range>261-4</page-range></nlm-citation>
</ref>
<ref id="B29">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Pounds]]></surname>
<given-names><![CDATA[J. A.]]></given-names>
</name>
<name>
<surname><![CDATA[Crump]]></surname>
<given-names><![CDATA[M. L.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Amphibian declines and climate disturbance: The case of the Golden Toad and the Harlequin Frog]]></article-title>
<source><![CDATA[Conservation Biology]]></source>
<year>1994</year>
<volume>8</volume>
<numero>1</numero>
<issue>1</issue>
<page-range>72-85</page-range></nlm-citation>
</ref>
<ref id="B30">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Pounds]]></surname>
<given-names><![CDATA[J. A.]]></given-names>
</name>
<name>
<surname><![CDATA[Fogden]]></surname>
<given-names><![CDATA[M. P.]]></given-names>
</name>
<name>
<surname><![CDATA[Campbell]]></surname>
<given-names><![CDATA[J. H.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Biological response to climate change on a tropical mountain]]></article-title>
<source><![CDATA[Nature]]></source>
<year>1999</year>
<volume>398</volume>
<page-range>611-5</page-range></nlm-citation>
</ref>
<ref id="B31">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Price]]></surname>
<given-names><![CDATA[P. W.]]></given-names>
</name>
<name>
<surname><![CDATA[Westoby]]></surname>
<given-names><![CDATA[M.]]></given-names>
</name>
<name>
<surname><![CDATA[Rice]]></surname>
<given-names><![CDATA[B.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Parasite-mediated competition: some predictions and tests]]></article-title>
<source><![CDATA[The American Naturalist]]></source>
<year>1988</year>
<volume>131</volume>
<numero>4</numero>
<issue>4</issue>
<page-range>544-55</page-range></nlm-citation>
</ref>
<ref id="B32">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Puschendorf]]></surname>
<given-names><![CDATA[R.]]></given-names>
</name>
<name>
<surname><![CDATA[Bolaños]]></surname>
<given-names><![CDATA[F.]]></given-names>
</name>
<name>
<surname><![CDATA[Chaves]]></surname>
<given-names><![CDATA[G.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[The amphibian chytrid fungus along an altitudinal transect before the first reported declines in Costa Rica]]></article-title>
<source><![CDATA[Biological Conservation]]></source>
<year>2006</year>
<volume>132</volume>
<page-range>136-42</page-range></nlm-citation>
</ref>
<ref id="B33">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Puschendorf]]></surname>
<given-names><![CDATA[R.]]></given-names>
</name>
<name>
<surname><![CDATA[Chaves]]></surname>
<given-names><![CDATA[G. A.]]></given-names>
</name>
<name>
<surname><![CDATA[Crawford]]></surname>
<given-names><![CDATA[A. J.]]></given-names>
</name>
<name>
<surname><![CDATA[Brooks]]></surname>
<given-names><![CDATA[D. R.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Eleutherodactylus ranoides (NCN). Dry forest population, refuge from decline?]]></article-title>
<source><![CDATA[Herpetological Review]]></source>
<year>2005</year>
<volume>36</volume>
<numero>1</numero>
<issue>1</issue>
<page-range>53</page-range></nlm-citation>
</ref>
<ref id="B34">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Rivera-Correa]]></surname>
<given-names><![CDATA[M.]]></given-names>
</name>
<name>
<surname><![CDATA[Jimenez-Rivillas]]></surname>
<given-names><![CDATA[C.]]></given-names>
</name>
<name>
<surname><![CDATA[Daza]]></surname>
<given-names><![CDATA[J. M.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Phylogenetic analysis of the Neotropical Pristimantis leptolophus species group (Anura: Craugastoridae): molecular approach and description of a new polymorphic species]]></article-title>
<source><![CDATA[Zootaxa]]></source>
<year>2017</year>
<volume>4242</volume>
<numero>2</numero>
<issue>2</issue>
<page-range>313-43</page-range></nlm-citation>
</ref>
<ref id="B35">
<nlm-citation citation-type="book">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Rodríguez-Salazar]]></surname>
<given-names><![CDATA[H.]]></given-names>
</name>
</person-group>
<source><![CDATA[Reserva Biológica Alberto Manuel Brenes: veinticinco años de conservación, investigación y bioalfabetización]]></source>
<year>2000</year>
<publisher-loc><![CDATA[San Ramón, Alajuela, Costa Rica ]]></publisher-loc>
<publisher-name><![CDATA[Universidad de Costa Rica]]></publisher-name>
</nlm-citation>
</ref>
<ref id="B36">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Ryan]]></surname>
<given-names><![CDATA[M. J.]]></given-names>
</name>
<name>
<surname><![CDATA[Fuller]]></surname>
<given-names><![CDATA[M. M.]]></given-names>
</name>
<name>
<surname><![CDATA[Scott]]></surname>
<given-names><![CDATA[N. J.]]></given-names>
</name>
<name>
<surname><![CDATA[Cook]]></surname>
<given-names><![CDATA[J. A.]]></given-names>
</name>
<name>
<surname><![CDATA[Poe]]></surname>
<given-names><![CDATA[S.]]></given-names>
</name>
<name>
<surname><![CDATA[Willink]]></surname>
<given-names><![CDATA[B.]]></given-names>
</name>
<name>
<surname><![CDATA[&#8230; Bolaños]]></surname>
<given-names><![CDATA[F.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Individualistic population responses of five frog species in two changing tropical environments over time]]></article-title>
<source><![CDATA[PLoS One]]></source>
<year>2014</year>
<volume>9</volume>
<numero>5</numero>
<issue>5</issue>
</nlm-citation>
</ref>
<ref id="B37">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Ryan]]></surname>
<given-names><![CDATA[F.]]></given-names>
</name>
<name>
<surname><![CDATA[Koerner]]></surname>
<given-names><![CDATA[S. E.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Too wet for frogs: changes in a tropical leaf-litter community coincide with La Niña]]></article-title>
<source><![CDATA[Ecosphere]]></source>
<year>2015</year>
<volume>6</volume>
<page-range>1-10</page-range></nlm-citation>
</ref>
<ref id="B38">
<nlm-citation citation-type="book">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Savage]]></surname>
<given-names><![CDATA[J. M.]]></given-names>
</name>
</person-group>
<source><![CDATA[The Amphibians and Reptiles of Costa Rica: A Herpetofauna between Two Continents, between Two Seas]]></source>
<year>2002</year>
<publisher-loc><![CDATA[Chicago ]]></publisher-loc>
<publisher-name><![CDATA[University of Chicago Press]]></publisher-name>
</nlm-citation>
</ref>
<ref id="B39">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Savage]]></surname>
<given-names><![CDATA[J. M.]]></given-names>
</name>
<name>
<surname><![CDATA[Kubicki]]></surname>
<given-names><![CDATA[B.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[A new species of fringe-limb frog, genus Ecnomiohyla (Anura: Hylidae), from the Atlantic slope of Costa Rica, Central America]]></article-title>
<source><![CDATA[Zootaxa]]></source>
<year>2010</year>
<volume>2719</volume>
<page-range>21-34</page-range></nlm-citation>
</ref>
<ref id="B40">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Scott]]></surname>
<given-names><![CDATA[N. J. Jr.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[The abundance and diversity of the herpetofauna of tropical forest litter]]></article-title>
<source><![CDATA[Biotropica]]></source>
<year>1976</year>
<volume>8</volume>
<page-range>41-58</page-range></nlm-citation>
</ref>
<ref id="B41">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Suzuki]]></surname>
<given-names><![CDATA[R.]]></given-names>
</name>
<name>
<surname><![CDATA[Shimodaira]]></surname>
<given-names><![CDATA[H.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Pvclust: an R package for assessing the uncertainty in hierarchical clustering]]></article-title>
<source><![CDATA[Bioinformatics]]></source>
<year>2006</year>
<volume>22</volume>
<numero>12</numero>
<issue>12</issue>
<page-range>1540-2</page-range></nlm-citation>
</ref>
<ref id="B42">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Voyles]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
<name>
<surname><![CDATA[Woodhams]]></surname>
<given-names><![CDATA[D. C.]]></given-names>
</name>
<name>
<surname><![CDATA[Saenz]]></surname>
<given-names><![CDATA[V.]]></given-names>
</name>
<name>
<surname><![CDATA[Byrne]]></surname>
<given-names><![CDATA[A. Q.]]></given-names>
</name>
<name>
<surname><![CDATA[Perez]]></surname>
<given-names><![CDATA[R.]]></given-names>
</name>
<name>
<surname><![CDATA[Rios-Sotelo]]></surname>
<given-names><![CDATA[G.]]></given-names>
</name>
<name>
<surname><![CDATA[&#8230; Richards-Zawacki]]></surname>
<given-names><![CDATA[C. L.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Shifts in disease dynamics in a tropical amphibian assemblage are not due to pathogen attenuation]]></article-title>
<source><![CDATA[Science]]></source>
<year>2018</year>
<volume>359</volume>
<numero>6383</numero>
<issue>6383</issue>
<page-range>1517-9</page-range></nlm-citation>
</ref>
<ref id="B43">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Whitfield]]></surname>
<given-names><![CDATA[S.]]></given-names>
</name>
<name>
<surname><![CDATA[Bell]]></surname>
<given-names><![CDATA[E.]]></given-names>
</name>
<name>
<surname><![CDATA[Philippi]]></surname>
<given-names><![CDATA[T.]]></given-names>
</name>
<name>
<surname><![CDATA[Sasa]]></surname>
<given-names><![CDATA[M.]]></given-names>
</name>
<name>
<surname><![CDATA[Bolaños]]></surname>
<given-names><![CDATA[F.]]></given-names>
</name>
<name>
<surname><![CDATA[Chaves]]></surname>
<given-names><![CDATA[G.]]></given-names>
</name>
<name>
<surname><![CDATA[&#8230; Donelly]]></surname>
<given-names><![CDATA[M.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Amphibian and reptile declines over 35 years at La Selva, Costa Rica]]></article-title>
<source><![CDATA[Proceedings of the National Academy of the United States of America]]></source>
<year>2007</year>
<volume>104</volume>
<page-range>8352-6</page-range></nlm-citation>
</ref>
<ref id="B44">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Woodhams]]></surname>
<given-names><![CDATA[D. C.]]></given-names>
</name>
<name>
<surname><![CDATA[Kilburn]]></surname>
<given-names><![CDATA[V. L.]]></given-names>
</name>
<name>
<surname><![CDATA[Reinert]]></surname>
<given-names><![CDATA[L. K.]]></given-names>
</name>
<name>
<surname><![CDATA[Voyles]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
<name>
<surname><![CDATA[Medina]]></surname>
<given-names><![CDATA[D.]]></given-names>
</name>
<name>
<surname><![CDATA[Ibáñez]]></surname>
<given-names><![CDATA[R.]]></given-names>
</name>
<name>
<surname><![CDATA[&#8230; Rollins-Smith]]></surname>
<given-names><![CDATA[L. A.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Chytridiomycosis and amphibian population declines continue to spread eastward in Panama]]></article-title>
<source><![CDATA[EcoHealth]]></source>
<year>2008</year>
<volume>5</volume>
<numero>3</numero>
<issue>3</issue>
<page-range>268-74</page-range></nlm-citation>
</ref>
</ref-list>
</back>
</article>
