<?xml version="1.0" encoding="ISO-8859-1"?><article xmlns:mml="http://www.w3.org/1998/Math/MathML" xmlns:xlink="http://www.w3.org/1999/xlink" xmlns:xsi="http://www.w3.org/2001/XMLSchema-instance">
<front>
<journal-meta>
<journal-id>0034-7744</journal-id>
<journal-title><![CDATA[Revista de Biología Tropical]]></journal-title>
<abbrev-journal-title><![CDATA[Rev. biol. trop]]></abbrev-journal-title>
<issn>0034-7744</issn>
<publisher>
<publisher-name><![CDATA[Universidad de Costa Rica]]></publisher-name>
</publisher>
</journal-meta>
<article-meta>
<article-id>S0034-77442019000200228</article-id>
<article-id pub-id-type="doi">10.15517/rbt.v67i2supl.37236</article-id>
<title-group>
<article-title xml:lang="en"><![CDATA[The skipper butterflies (Lepidoptera: Hesperiidae) of the Reserva Ecológica Leonelo Oviedo, San José, Costa Rica]]></article-title>
</title-group>
<contrib-group>
<contrib contrib-type="author">
<name>
<surname><![CDATA[Murillo-Hiller]]></surname>
<given-names><![CDATA[Luis Ricardo]]></given-names>
</name>
<xref ref-type="aff" rid="Aff"/>
<xref ref-type="aff" rid="Aaf"/>
<xref ref-type="aff" rid="A a"/>
<xref ref-type="aff" rid="A3"/>
</contrib>
<contrib contrib-type="author">
<name>
<surname><![CDATA[Segura-Bermúdez]]></surname>
<given-names><![CDATA[Oscar Antonio]]></given-names>
</name>
<xref ref-type="aff" rid="Aff"/>
</contrib>
<contrib contrib-type="author">
<name>
<surname><![CDATA[Barquero]]></surname>
<given-names><![CDATA[Juan Diego]]></given-names>
</name>
<xref ref-type="aff" rid="Aff"/>
</contrib>
<contrib contrib-type="author">
<name>
<surname><![CDATA[Bolaños]]></surname>
<given-names><![CDATA[Federico]]></given-names>
</name>
<xref ref-type="aff" rid="Aff"/>
<xref ref-type="aff" rid="Aaf"/>
</contrib>
</contrib-group>
<aff id="Af1">
<institution><![CDATA[,Universidad de Costa Rica  ]]></institution>
<addr-line><![CDATA[ San José]]></addr-line>
<country>Costa Rica</country>
</aff>
<aff id="Af2">
<institution><![CDATA[,Universidad de Costa Rica  ]]></institution>
<addr-line><![CDATA[ ]]></addr-line>
<country>Costa Rica</country>
</aff>
<aff id="Af3">
<institution><![CDATA[,Universidad de Costa Rica  ]]></institution>
<addr-line><![CDATA[ ]]></addr-line>
<country>Costa Rica</country>
</aff>
<pub-date pub-type="pub">
<day>00</day>
<month>04</month>
<year>2019</year>
</pub-date>
<pub-date pub-type="epub">
<day>00</day>
<month>04</month>
<year>2019</year>
</pub-date>
<volume>67</volume>
<numero>2</numero>
<fpage>228</fpage>
<lpage>248</lpage>
<copyright-statement/>
<copyright-year/>
<self-uri xlink:href="http://www.scielo.sa.cr/scielo.php?script=sci_arttext&amp;pid=S0034-77442019000200228&amp;lng=en&amp;nrm=iso"></self-uri><self-uri xlink:href="http://www.scielo.sa.cr/scielo.php?script=sci_abstract&amp;pid=S0034-77442019000200228&amp;lng=en&amp;nrm=iso"></self-uri><self-uri xlink:href="http://www.scielo.sa.cr/scielo.php?script=sci_pdf&amp;pid=S0034-77442019000200228&amp;lng=en&amp;nrm=iso"></self-uri><abstract abstract-type="short" xml:lang="en"><p><![CDATA[Abstract Hesperiidae is one of the most diverse families of butterflies in Costa Rica, with approximately 486 species. Even so, there are few butterfly lists where this group has been included. In this paper, we present information on seasonality, abundance and natural history features of this family for the Leonelo Oviedo Ecological Reserve (RELO), a 2 ha forest embedded in an urban matrix. Over the course of two years, a monthly sampling was carried out on a 270 m trail across the Reserve from 08:00 to 12:00, collecting all the individuals located within 5 m on each side of the trail. To better represent the richness, individuals were also randomly collected for more than ten years, but the butterflies collected in this way were not included in the statistical analysis. Photographs were taken of all the species in order to provide an identification guide. For the cryptic species, drawings and dissections of the genitalia were made. For the community indexes we used Microsoft Excel and the Shannon index with base two logarithm. For the summary of the monthly data analysis were done according to dry and wet season. For a comparison of richness and abundance we did a g-test to evaluate if there are differences between seasons; however, with the use of the R package vegan a hierarchical cluster analysis was done using the Jaccard index with Wards minimum variance agglomerative method. With R package pvclust the uncertainty of the clusters based on a bootstrap with 10 000 iterations. 423 individuals of 49 species were included in the statistical analysis, from a total of 435 individuals of 58 species. A tendency to greater richness and abundance of skippers was found during the dry season. Through the cluster analysis, it was possible to determine that in relation to the diversity of skippers, both wet seasons are grouped significantly (P = 0.05). The dry seasons are also grouped significantly (P = 0.05). The reserve has connectivity with other green areas via a stream. During the wet season, plant growth increases connectivity, which could lead to the entry of new individuals of different species that are not permanent residents of RELO and establish small populations, increasing the richness and abundance of species. This added to the variation in the occurrence of some species of butterflies in response to seasonal variations and differences in the availability of resources in different seasons explains the grouping of species between seasons.]]></p></abstract>
<abstract abstract-type="short" xml:lang="es"><p><![CDATA[Resumen Con aproximadamente 486 especies, Hesperiidae es una de las familias más diversas de mariposas de Costa Rica. A pesar de eso, en pocas listas de especies locales son incluidas. En este trabajo, presentamos información sobre la estacionalidad, abundancia e historia natural de los hespéridos de la Reserva Ecológica Leonelo Oviedo. Ésta reserva se encuentra dentro de una matriz urbanizada y por un período de dos años se realizaron muestreos mensuales a lo largo de un sendero de 270 m que la atraviesa. Las recolectas se llevaron a cabo desde las 08:00 hasta las 12:00, atrapando todos los individuos encontrados a cada lado del sendero a una distancia de hasta 5 m. Para representar mejor la riqueza de especies, se siguió recolectando individuos de manera aleatoria por más de diez años posteriores al muestreo inicial. Consecuentemente, estos individuos no fueron incluidos en el análisis estadístico realizado. Se tomaron fotografías de todas las especies obtenidas en total, para proporcionar una guía de identificación de especies. Ademas, para las especies crípticas se realizaron disecciones e ilustraciones de los genitales masculinos. Para el análisis de diversidad se utilizó el índice de Shannon con logaritmo en base dos y analizando los datos por separado de las estación lluviosa y seca. Para la comparación de riqueza y abundancia se realizó una prueba G y se obtuvo que hay mayor diversidad y abundancia durante la estación seca. Se utilizó el paquete vegan R para un análisis de conglomerados de Jaccard con un indice Ward de mínima varianza. Con el Paquete R pvclust la incertidumbre de los conglomerados se realizó con un bootstrap de 10 000 interacciones. 423 individuos de 49 especies fueron incluidas en el análisis estadístico, de un total de 435 individuos de 58 especies tratadas en este trabajo. Con el análisis de conglomerados fue posible determinar que la diversidad de Hesperiidae en las dos estaciones lluviosas analizadas se agrupan significativamente (P = 0.05). Lo mismo que sucede con las estaciones secas (P = 0.05). La Reserva presenta conectividad con otras áreas boscosas a través de una quebrada. Durante la estación lluviosa, el crecimiento vegetativo aumenta el paso de individuos lo que puede propiciar la llegada de nuevas especies que no necesariamente son residentes permanentes de la Reserva y establecerse en pequeñas poblaciones que incrementan la diversidad y abundancia temporalmente, Lo anterior, mas las variaciones estacionales en la disponibilidad de recursos explican la similitud de la composición de especies entre cada una de las estaciones.]]></p></abstract>
<kwd-group>
<kwd lng="en"><![CDATA[&#946;-biodiversity]]></kwd>
<kwd lng="en"><![CDATA[conservation]]></kwd>
<kwd lng="en"><![CDATA[species richness]]></kwd>
<kwd lng="en"><![CDATA[urban forest reserve]]></kwd>
<kwd lng="en"><![CDATA[bio-indicators]]></kwd>
<kwd lng="es"><![CDATA[biodiversidad &#946;]]></kwd>
<kwd lng="es"><![CDATA[conservación]]></kwd>
<kwd lng="es"><![CDATA[riqueza de especies]]></kwd>
<kwd lng="es"><![CDATA[reserva biológica urbana]]></kwd>
<kwd lng="es"><![CDATA[bioindicadores de calidad de hábitat]]></kwd>
</kwd-group>
</article-meta>
</front><back>
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