The collecting trips by Ørsted and Kröyer in Central and South America resulted in a series of papers by Grube, the Annulata örstediana which contained the proposal for seven genera and descriptions for 84 species. There are some problems dealing with the correct number of contributions, their publication dates, the correct citation for the authors for each species name and sometimes even for the type locality. In order to improve the current situation, we have consulted the original publications together with the corresponding collections. Our results indicate that there were only three parts in the series which were published in the journal, and repagination in reprints sometimes caused some further confusion. Additionally, we provided some review comments on the species current status and we added a short nomenclatural note on Hemipodus (Polychaeta: Glyceridae) trying to clarify the correct generic name and type species.
Los viajes de recolecta por Ørsted y Kröyer en Centroamérica y Sudamérica resultaron en una serie de notas por Grube, los Annulata örstediana que contienen la propuesta de siete géneros y 84 especies. Hay algunos problemas en relación con el número correcto de contribuciones, sus fechas de publicación, la citación correcta de los autores para cada nombre específico y a veces incluso para la localidad tipo. Para mejorar la situación, hemos consultado las publicaciones originales junto con las colecciones correspondientes. Nuestros resultados indican que hubo tres partes en la serie, que fueron publicadas en la revista y cuyas páginas fueron re-enumeradas para las separatas lo que a veces ocasionó confusiones adicionales. Además, brindamos algunos comentarios para revisar el estado de cada especie y agregamos una breve nota nomenclatural sobre Hemipodus (Polychaeta: Glyceridae) para aclarar el nombre correcto del género y la especie tipo.
Palabras clave: taxonomía, nomenclatura, invertebrados marinos, hirudineos, sipúnculos. The series of papers entitled Annulata örstediana described species collected by Anders S. Ørsted and Henrik Kröyer in the Caribbean, Central America and South America. Adolph-Eduard Grube provided standardized descriptions and indicated the corresponding co-author for each name, but the correct authors for them have been elusive and, at the same time, the correct publication dates have been misunderstood. Most species were polychaetes but some hirudineans and sipunculans were also included; seven genera were proposed and 84 species were newly described.
Some of the species were studied by Augener (1925) and further comments were made by Wolff & Petersen (1991). In the latter contribution, the authors tried to clarify the publication dates for the series. However, two problems remain for correct citation of the species. First, despite the fact that many species were introduced with two authors (‘Gr. Örsd.’ or ‘Gr. Kr.’), most species have been attributed to Grube alone. This contradicts Recommendation 51E of the Zoological Code (ICZN 1999), but probably because it is just a recommendation, it has not been widely followed. We think this should be modified such that the correct authors are acknowledged and some examples have already been introduced (Hartwich 1993). Second, because the parts were compiled and distributed as collected reprints with different pagination, there seems to be confusion about how many parts were issued. Hartman (1951a:101) indicated three parts, whereas Wolff & Petersen (1991:680) listed four. To clarify this issue, the corresponding volumes of the journal (Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening) have been thoroughly rechecked and the full citations are included to avoid confusion in the future. Furthermore, a list of all taxa newly described in this series is provided in order to improve the recognition of their correct authors together with some notes on the current status.
Recommended citations
(Publications available in http://www. archive.org/stream/annulatarstedi00grub/annulatarstedi00grub_djvu.txt)
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Grube, A.E. 1857. Annulata örstediana. Enumeratio Annulatorum, quae in itinere per Indiam occidentalem et Americam centralem annis 1845-1848 suscepto legit cl. A.S. Örsted, adjectis speciebus nonnullis a cl. H. Kröyero in itinere ad Americam meridionalem collectis. [1. Familia Aphroditea – F. Euniceae]. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening for 1856: 44-62.
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Grube, A.E. 1858. Annulata örstediana. Enumeratio Annulatorum, quae in itinere per Indiam occidentalem et Americam centralem annis 1845-1848 suscepto legit cl. A.S. Örsted, adjectis speciebus nonnullis a cl. H. Kröyero in itinere ad Americam meridionalem collectis, (Fortsættelse [continued]) [2. Familia Euniceae F. Syllidea]. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening for 1857: 158-186. ]]>
Grube, A.E. 1859. Annulata örstediana. Enumeratio Annulatorum, quae in itinere per Indiam occidentalem et Americam centralem annis 1845-1848 suscepto legit cl. A.S. Örsted, adjectis speciebus nonnullis a cl. H. Kröyero in itinere ad Americam meridionalem collectis, (Slutning [conclusion]) [3. Familia Ariciaea F. Sipunculacea and Index]. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening for 1858: 105-120.
Annotated taxonomic list
The following list includes only the newly described taxa; they are presented for each family (except in Hirudinea and Sipuncula) and all in ]]>
Nomenclator ]]>
in http:// uio.mbl.edu/NomenclatorZoologicus/.
Polychaeta
Amphinomidae ]]>
Amphinome
1. A. carnea Grube & Ørsted in Grube, 1857: 52, St. Croix. The status of this species was regarded as uncertain by Baird (1870:219). Hartman (1959:128) regarded it as ]]>
2. A. paupera Grube & Kröyer in Grube, 1857: 52-53, Valparaiso, ZMB 2785 (3 syntypes). Augener (1922:172) regarded this species as a subspecies in E. complanata; Hartman (1959:129) referred it to Eurythoe as E. ]]>
. WesenbergLund (1962:38) transferred this species to Pareurythoe.
Notopygos
3. N. ]]>
Grube & Ørsted in Grube, 1857: 53, Puntarenas, Costa Rica, ZMUCPOL-183 (3 syntypes), ZMB 3331 (lost). The species was redescribed (Monro 1933a:10) and the name is available (Hartman 1940:207, 1959:136).
Chrysopetalidae ]]>
4. Palmyra(?) elongata Grube & Ørsted in Grube, 1857: 51-52, St. Croix, ZMUC- POL-251 (holotype). Augener (1925:5) regarded this species as belonging in Bhawania and Hartman (1959:127) thought it might be identical to B. goodei ]]>
Cirratulidae
Cirratulus
5. C. caribous Grube & Ørsted in Grube, 1859: 106, St. Croix, 3 fathoms. Hartman (1959:403) regarded this species as indeterminable. ]]>
6. C. punctatus Grube & Ørsted in Grube, 1859: 107, St. Croix, Christiansted, Virgin Islands, 3 fathoms. Monro (1933b:264) recorded this species from Southern Florida. Hartman (1959:405) transferred it to Cirriformia. The species has been redescribed and transferred to Timarete Kinberg, 1866 (Çinar 2007:757). ]]>
Dorvilleidae
7. Anisoceras Grube & Ørsted in Grube, 1857: 60. Pre-occupied by Anisoceras Hope, 1840, ]]>
Papilliodorvillea as a replacement name, with A. rubra (see below) as the type species. The presence of a nuchal papilla was rejected as a generic, diagnostic feature, and the genus was considered a junior synonym of Dorvillea by Jumars (1974:114). It should be noted that Dorvillea Parfitt, 1866 is a junior homonym of Dorvillea Gray, ]]>
8. A. bioculata Grube & Ørsted in Grube, 1857: 62, Puntarenas, Costa Rica. Illustrated by Wolff & Petersen (1991:Fig. 2A-F). Hartman (1959:348) regarded this species as a member of Dorvillea Parfitt, ]]>
9. A. rubra Grube & Ørsted in Grube, 1857: 60-61, St. Croix, 30 fathoms. This species was regarded as belonging in Dorvillea (Hartman 1951b:66, 1959:348). It was later designated as the type species of Papilliodorvillea Pettibone, ]]>
10. A. vittata Grube & Ørsted in Grube, 1857: 61-62, Puntarenas, Costa Rica. Illustrated by Wolff & Petersen (1991:Pl. 2, Fig. 7). According to Hartman (1959:348), the species belongs in Dorvillea and this was confirmed by Pettibone (1961:182).
Eunicidae Eunice
11. E. cariboea Grube & ]]>
in Grube, 1857: 57, St. Croix, Christiansted, Virgin Islands, ZMUC-POL-297 (lectotype, designated by K. Fauchald), ZMUC-POL-298 (1 paralectotype), ZMUC-POL-299 (1 paralectotype = holotype of E. excariboea Fauchald, 1992), ZMUC-POL-300 (1 paralectotype = paratype of E. excariboea), ZMB 4004 (3 syntypes and 3 anterior fragments). Monro (1933a:63) recorded this species from the Eastern Tropical Pacific. It was redescribed by ]]>
12. E. filamentosa Grube & Ørsted in Grube, 1857: 56, St. Croix, ZMUC-POL-303 (lectotype, designated by K. Fauchald), ZMUC-POL-304 (1 paralectotype = Palola sp. fide Fauchald 1992), ZMB 1855 (1 paralectotype). Augener (1925) reexamined this species and found that it contained a specimen that he thought belonged to
E. gagzoi Augener, 1922 or E. siciliensis Grube, 1840; this specimen has subsequently been referred to Palola (K. Fauchald, pers. comm.). Monro (1933a:65) recorded it from the Eastern Tropical Pacific. The species was redescribed and restricted by Carrera-Parra & Salazar-Vallejo (2011:53-56, Fig. 1). 13. E. lucei Grube & ]]>
in Grube, 1857: 57-58, Puntarenas, Costa Rica, ZMUCPOL-305 (lectotype, designated by K. Fauchald), ZMUC-POL-306 (1 paralectotype), ZMB 2019 (holotype). This species was regarded as indeterminable by Hartman (1959:312) but it was redescribed by Fauchald (1992:201); he also selected a lectotype from among the Copenhagen specimens, but this does not appear to be valid since the holotype is in Berlin (Hartwich 1993:114). 14.
E. punctata Grube & Ørsted in Grube, 1857: 59-60, St. Thomas, ZMUC-POL-307 (holotype = holotype of E. binominata de Quatrefages, 1866). According to Ehlers (1887:85), Augener (1906:132) and Hartman (1959:314), this is a junior homonym of E. punctata (Risso, 1826); de Quatrefages (1866:327) proposed as a replacement name, but the single specimen was thought ]]>
species inquerenda. The specimen is, however, present in the collection of the Zoological Museum in Copenhagen and should be regarded as the holotype of E. binominata de Quatrefages, 1866. 15. E. quadrioculata ]]>
in Grube, 1857: 60, Puntarenas, Costa Rica. Fauchald (1992:284) commented that this species was described on the basis of an illustration and that de Quatrefages (1866:337) referred it to Marphysa; however, the latter author regarded the species as being deprived of branchiae and probably belonging in Lumbrineridae. 16. E. rubra ]]>
in Grube, 1857: 59, St. Thomas, ZMUC-POL-308 (holotype). Redescribed by Ehlers (1887:87), Augener (1925:9), Hartman (1951b:55) and Fauchald (1992:291). 17. E. splendida Grube & Kröyer in Grube, 1857: 58, Valparaiso, ZMUC-POL-309 (holotype). ]]>
18. E. violacea Grube & Ørsted in Grube, 1857: 55, Puntarenas, Costa Rica. Fauchald (1992:335) regarded this species as indeterminable.
Flabelligeridae Siphonostomum
19. S. cariboum Grube & Ørsted in Grube, 1859: 108-109, St. Croix, ZMUC-POL-331 (holotype). Ehlers (1887:158) recorded ]]>
Semiodera. 20. S. cingulatum Grube & Kröyer in Grube, 1859: 108, Rio de Janeiro.
Glyceridae ]]>
Glycera
21. G. cirrata Grube, 1858: 176, Rio de Janeiro, St. Croix, ZMUC-POL-392 (1 syntype). According to Augener (1925:10) the specimen present in the Zoological Museum in Copenhagen, which is from St. Croix, Virgin Islands, is not the one that ]]>
G. tesselata Grube, 1863, described from the Mediterranean Sea. Hartman (1950:64, 1959:289) regarded this species as close to G. americana Leidy, 1855 described from Rhode Island. Böggemann (2002:23) indicated that there were two lots, one from the Virgin Islands, which he regards as belonging to G. americana, whereas the Brazilian materials belong to
G. brevicirris Grube, 1870, which has an uncertain type locality, probably the Indian Ocean. 22. G. papillosa Grube, 1858: 176-177, Valparaíso. Hartman (1940:248) followed Augener (1922:203) and based on a specimen from Perú, regarded the species as valid. It was briefly characterized by WesenbergLund (1962:103); however, because there is no type material, Böggemann (2002:28) regarded it as nomen dubium. ]]>
Hemipodus de Quatrefages, 1866 or Hemipodia de Quatrefages in Kinberg, 1866? Böggemann (2002:79) proposed Hemipodia Kinberg, 1865 as a senior synonym of Hemipodus de Quatrefages, 1866, and designated Hemipodia simplex as the type ]]>
nomina nuda. Being aware of this, Kinberg ]]>
Hemipodia Quatref.” and referred to the first of the earlier proposals by de Quatrefages, which did not include descriptions or illustrations. Consequently, this new generic name is not a different genus but rather an orthographic correction, which was clearly indicated by Kinberg himself and emphasized by Grube (1870:66); we think that if this replacement name is really older than the Latin proposal of Hemipodus de Quatrefages (1866:194), then it should be cited as Hemipodia de ]]>
H. patagonica as the only species in Hemipodia, and according to article 68.3 (ICZN 1999:71), this is the type species by monotypy. However, de Quatrefages (1866:194) did not introduce the five species in alphabetical order and the first species was H. ]]>
de Quatrefages, 1866, which was subsequently designated by Hartman (1940:79) as the type species of Hemipodus. It is enigmatic why Böggemann thought that a new type species was needed, since there are two type species for these two generic names, but according to article 67.1.2 (ICZN 1999:66), H. simplex cannot be the type species, despite the fact that it is the senior synonym of H. patagonica Kinberg, 1866.
23. H. simplex Grube, 1858: 177-178, Valparaiso, Callao. This species was actually described within Glycera but was introduced as “H. simplex” due to a typographic error (similarly, in his comments on the preceding species, Glycera
papillosa, Grube referred to G. capitata as H. capitata). It was briefly characterized by Wesen- berg-Lund (1962:104) and redescribed by Böggemann (2002:79-81, Figs. 127-129).
Hesionidae
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Oxydromus Grube, 1855. Pre-occupied by Oxydromus Schlegel, 1854 (Pleijel 1998:161; see Ophiodromus).
24. O. flaccidus Grube & Ørsted in Grube, ]]>
nomendubium because the type material was probably lost. 25. O. longisetis Grube & Ørsted in Grube, 1858172-173, St. Croix. Pleijel (1998:162) regarded this species as nomen ]]>
dubium because the type material was probably lost.
Lumbrineridae Lumbrineris
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26. L. brasiliensis Grube & Kröyer in Grube, 1858: 159, Rio de Janeiro, ZMB 5592 (3 syntypes). Resembles species in Scoletoma, but the genus has not been revised (L.F. Carrera-Parra, 2011, com. pers.) 27. L. longissima Grube & ]]>
in Grube, 1858: 158, Valparaiso. Hartman (1959:329) regarded this species as an oenonid, probably Arabella iricolor (Montagu, 1804), which was described from England.
Nephtyidae ]]>
Nephtys
28. N. imbricata Grube & Kröyer in Grube, 1858: 168, Valparaiso, ZMUC-POL-1471 (1 syntype), ZMB 1881 (1 syntype). Hartman (1959:283) regarded this species as a probable synonym of N. squamosa ]]>
Nereididae Heteronereis
29.
H. caudipunctata Grube & Ørsted in Grube, 1858: 166-167, St. Croix, 20 fathoms, ZMUC-POL-1494 (holotype). Hartman (1959:242) transferred this species to Nereis. 30. H. pannosa Grube & Kröyer in Grube, 1858: 167-168, Callao. ]]>
Nereilepas
31. N. callaona ]]>
in Grube, 1858: 165-166, Callao, ZMUC-POL-1495 (holotype). Hartman (1939a:11, 1940:227, 1959:251) transferred this species to Nereis. It was recorded and briefly redescribed by Hartmann-Schröder (1960:19, 1962:107) and by Wesenberg-Lund (1962:75). 32. N. variegata Grube & ]]>
in Grube, 1858: 164-165, Valparaíso, ZMUCPOL-1496 (holotype). Augener (1927:48) regarded this species as a member of Perinereis and Monro (1933a:45, Hartman 1959:251, Wesenberg-Lund 1962:83) transferred it to Pseudonereis.
Nereis
33. N. debilis Grube & Ørsted in Grube, 1858: 164, no type locality indicated. Hartman (1959:257) transferred this species to Ceratonereis. ]]>
34. N. krebsii Grube & Ørsted in Grube, 1858: 161, St. Croix, ZMUC-POL-1482 (holotype). This species should possibly be transferred to either Pseudonereis or Perinereis (Augener 1925:7). Hartman (1959:261) regarded it as indeterminable. ]]>
35. N. marginata Grube & Ørsted in Grube, 1858: 160-161, St. John, ZMUC-POL-1483 holotype). Hartman (1959:264) regarded this species as indeterminable. 36. N. puncturata Grube & Ørsted in Grube, 1858: 163, Valparaiso and St. Croix (Grube ]]>
fide Hartwich 1993:130). Hartman (1959:269) regarded this species as indeterminable. 37. N. rigida Grube & Ørsted in Grube, 1858: 162, Puntarenas, Costa Rica. Redescribed by Augener (1906:149) on the basis of ]]>
38. N. riisei Grube & Ørsted in Grube, 1858: 162-163, Puntarenas, Costa Rica, ZMUCPOL-1487 (1 syntype). This species was redescribed by Augener (1925:6), Hartman (1940:221) and Monro (1933a:43) and was later recorded for the Caribbean fauna (Augener 1927:47, Monro 1933b:256, Hartman 1942:99, 1944:17, 1951b:46, Renaud 1956:14). However, N. glandulata ]]>
N. riisei. The syntype in the Zoological Museum in Copenhagen is the smaller of the two specimens mentioned in the original description; the whereabouts of the larger specimen is unknown. 39.
N. vallata Grube & Kröyer in Grube, 1858: 159-160, Valparaiso, ZMUC-POL-1489 (1 syntype), ZMUC-POL-1490 (1 syntype), ZMUC-POL-1491 (1 syntype), ZMUCPOL-1492 (1 syntype), ZMUC-POL-1493 (1 syntype). Hartman (1959:272) transferred this species to Perinereis, and it was redescribed by Wesenberg-Lund (1962:79) and Bertrán (1980:84). According to the original description only four specimens were found, but the type material contains five syntypes, which were placed in separate vials ]]>
Onuphidae Diopatra
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40. D. brevicirris Grube & Kröyer in Grube, 1857: 55, Madeira. 41. D. rhizophorae Grube & Ørsted in Grube, 1857: 54, Realejo, Nicaragua, ZMUCPOL-1507 (3 syntypes).
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Opheliidae Ophelia
42. O. appendiculata Grube & Kröyer in Grube, 1859: 107, Valparaiso. Hartman (1959:432) transferred this species to Ophelina ]]>
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45. P. callaona Grube & Kröyer in Grube, 1858: 170-171, Callao. 46. P. flavescens Ørsted in Grube, 1858: 172, Puntarenas, Costa Rica, ZMUC-POL-1563 (lectotype). ZMUC-POL-1563 is not a specimen, but rather ]]>
species inquerenda. 47. P. longicirris Grube & Ørsted in Grube, 1858: 171-172, Realejo, Nicaragua. Hartman (1959:162) transferred this species to Kinbergia, which ]]>
nomen dubium. 48. P. macrophthalma Grube & Ørsted in Grube, 1858: 169, Realejo, Nicaragua. Hartman (1959:162) transferred this species to Kinbergia, which Pleijel (1991:257) regarded as a nomen ]]>
Pleijel also considered this species to be a nomen dubium. 49. P. puntarenae Grube & Ørsted in Grube, 1858: 169-170, Puntarenas, Costa Rica. Wesenberg-Lund (1962:44) regarded the species as
species inquerenda.
Pisionidae
50. Pisione Grube, 1858: 174-175. 51.
P. örstedii Grube, 1858: 175, Valparaiso, ZMUC-POL-1605 (1 syntype), ZMB 3834 (2 syntypes and slide). Hartman (1939b:91, 1959:124) redescribed the species and corrected the epithet to oerstedii, whereas Hartmann-Schröder (1962:70) and Wesenberg-Lund (1962:39) omitted the last letter.
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Polynoidae
52. Hermenia Grube & Ørsted in Grube, 1857:44. 53. H. verruculosa Grube ]]>
in Grube, 1857: 44-45, St. John, ZMUC-POL-1667 (holotype). The generic and specific names are valid (Augener 1925:4). Fauvel (1953:4) provided additional information and Ebbs (1966:500) redescribed the species. Hartman (1939a:4) recorded it from the Eastern Tropical Pacific but this might be a different species.
Polynoe
54. P. clavata Grube & Ørsted in Grube, 1857: 47-48, Christiansted and St. Croix. Hartman (1959:99) regarded this species as the same as Halosydna muelleri (see below) or H. parva Kinberg, 1855, described from Chile. ]]>
55. P. crucis Grube & Ørsted in Grube, 1857: 47, Christiansted, Virgin Islands, ZMUCPOL-1300 (2 syntypes). Transferred to Harmothoe by Augener (1925:4) and redescribed by Salazar-Silva (2010). 56. P. exanthema Grube & ]]>
in Grube, 1857: 46, Valparaiso, ZMUC-POL-1671 (holotype). Hartman (1939b:52, 1959:100, followed by Wesenberg-Lund 1962:23) transferred this species to Harmothoe, but Hartmann-Schröder (1965:67) transferred it to Lagisca Malmgren, 1865. 57. P. marginata Grube & Kröyer in Grube, 1857: ]]>
58. P. mülleri Grube & Kröyer in Grube, 1857: 48, Valparaíso. Hartman (1959:104) transferred this species to Halosydna, perhaps as a synonym of H. parva. 59. P. nigrovittata Grube ]]>
in Grube, 1857: 50. Hartman (1959:104) regarded this species as indeterminable. 60. P. punctulata Grube & Kröyer in Grube, 1857: 49, Rio de Janeiro. Hartman (1959:105) transferred this species to Halosydnella. ]]>
61. P. savignyi Grube & Kröyer in Grube, 1857: 45, Callao. Hartman (1959:106) transferred this species to Lepidonotus. 62. P. tenuisetis Grube & Kröyer in Grube, 1857: ]]>
63. P. tomentosa Grube & Ørsted in Grube, 1857: 45, Puntarenas, Costa Rica, ZMUCPOL-1317 (holotype). Hartman (1959:106) transferred this species to Lepidonotus. ]]>
64. P. variegata Grube & Kröyer in Grube, 1857: 49-50, Madeira. Hartman (1959:107) regarded this species as a junior synonym of P. scolopendrina Savigny, 1822, described from the Mediterranean Sea according to Hartman, but it was actually described from the Atlantic French coasts, which Savigny called “l’Océan”. ]]>
Sabellidae Sabella
65. S. brevicollaris Grube & Ørsted in Grube, ]]>
Hypsicomus Grube, 1870. 66. S. pacifici Grube & Ørsted in Grube, 1859: 113, Puntarenas, Costa Rica, ZMUCPOL-1985 (holotype). Hartman (1959:562) regarded this species as indeterminable. ]]>
Sigalionidae
67. Sigalion(?) pergamentaceum Grube & Ørsted in Grube, 1857: 50-51, St. Croix. Hartman (1959:119) regarded this species as questionable.
Syllidae
68. Amblyosyllis Grube & Ørsted in Grube, 1858: 186. ]]>
69. A. rhombeata Grube & Ørsted in Grube, 1858: 186, St. Croix. Illustrated by Wolff and Petersen (1991:Fig. 3A).
Syllis
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70. S. brachycirris Grube & Ørsted in Grube, 1858: 179, St. Croix, ZMUC-POL-920 (holotype). This species was regarded as identical to S. gracilis Grube, 1840, described from the Mediterranean Sea (Augener 1925:7; Hartman 1959:224). 71. S. breviarticulata Grube & ]]>
in Grube, 1858: 179-180, St. Croix. Hartman (1959:224) regarded this species as indeterminable. 72. S. latifrons Grube & Kröyer in Grube, 1858: 178-179, Callao. 73.
S. longesegmentata Grube & Ørsted in Grube, 1858: 184-185, St. Croix. 74. S. macroceras Grube & Ørsted in Grube, 1858: 185-186, St. Croix, Christiansted, Virgin Islands. Hartman (1959:227) regarded this species as indeterminable. 75. S. obscura Grube & Ørsted in Grube, 1858: 181. Hartman (1959:228) regarded this species as indeterminable. 76. S. rubra Grube & Ørsted in Grube, ]]>
Trypanosylliszebra Grube, 1860, described from France, and the other syntype (no longer present in the collection of the Zoological Museum in Copenhagen) was referred to Typosyllis corallicoloides Augener, 1922. This synonymy was repeated by Hartman (1959:229). 77. S. streptocephala Grube & Ørsted in Grube, 1858: 181-182, St. Croix, ZMUCPOL-922 holotype). Identical with Haplosyllis spongicola (Grube, 1855), described from the Mediterranean Sea (Augener 1925:8; Hartman 1959:229). ]]>
78. S. valida Grube & Kröyer in Grube, 1858: 183-184, Valparaíso, ZMUC-POL-881 (lectotype, designated by F. Licher in 1996), ZMUC-POL-890 (1 paralectotype). Hartman (1959:231) regarded this species as indeterminable and WesenbergLund (1962:58) considered it as a species inquerenda. 79.
S. zonata Grube & Ørsted in Grube, 1858182-183, St. Croix. Hartman (1959:231) regarded this species as indeterminable.
Terebellidae
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80. Sabellina Dujardin, 1839. This genus is questionable according to Hartman (1959:566) and was regarded as indeterminable by Fauchald (1977:140); both regarded it as a member of Sabellidae, but the only illustrated species, S. brachycera, resembles a juvenile, probably abranchiate terebellid (Dujardin 1839, Pl. 7, Figs. 6-8). 81.
S. longicauda Grube & Ørsted in Grube, 1859: 111, St. Croix, Christiansted, Virgin Islands. Hartman (1959:566) regarded this species as indeterminable.
Terebella
82.
T. alata Grube & Ørsted in Grube, 1859: 110, Puntarenas, Costa Rica. Hartman (1959:523) regarded this species as questionable. 83. T. comata Grube & Kröyer in Grube, 1859: 109, Valparaiso, Chile, Puntarenas, ZMUC-POL-2151 ]]>
Thelepus Leuckart, 1849. 84. T. frondosa Grube & Ørsted in Grube, 1859: 110-111, Puntarenas, Costa Rica, ZMUC-POL-2152 (holotype). Hartman (1959:524) regarded this species as questionable. ]]>
Hirudinea Aulacostomum
85. A. costaricense Grube ]]>
in Grube, 1859: 114, Cartago? 86. Centropygos Grube & Ørsted in Grube, 1859: 115. 87. C. joseensis Grube & Ørsted
in Grube, 1859: 115, San José. Redescribed by Badham (1923).
Clepsine
88. C. triserialis Grube ]]>
Sipuncula
89. Dendrostomum Grube & Ørsted in Grube, ]]>
90. D. alutaceum Grube & Ørsted in Grube, 1859: 118, St. Croix. According to WoRMS, this genus is a junior synonym of Themiste and referred to this species as T. (T.) alutacea. ]]>
Phascolosoma
91. P. antillarum Grube & Ørsted in Grube, 1859: 117-118, Puntarenas, Costa Rica, St. Croix. According to WoRMS, this species has been transferred to ]]>
Antillesoma as A. antillarum. 92. P. puntarenae Grube & Ørsted in Grube, 1859: 117, Puntarenas, Costa Rica, ZMUC-SIP-001 (2 syntypes). According to WoRMS, this species is a junior synonym of P. nigrescens (Keferstein, 1865). ]]>
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*Correspondencia: Sergio I. Salazar-Vallejo: Depto. de Ecología Acuática, El Colegio de la Frontera Sur, Chetumal, México; ssalazar@ecosur.mx Danny Eibye-Jacobsen: Zoological Museum, Natural History Museum of Denmark, Copenhagen, Denmark; dejacobsen@snm.ku.dk 1. Depto. de Ecología Acuática, El Colegio de la Frontera Sur, Chetumal, México; ssalazar@ecosur.mx 2. Zoological Museum, Natural History Museum of Denmark, Copenhagen, Denmark; dejacobsen@snm.ku.dk
Received 29-VII-2011. Corrected 25-I-2012. Accepted 22-II-2012.
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